Exocelina

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Dytiscidae
Subfamilia: Copelatinae
Genus: Exocelina
Species (214):
Directory A·B·C·D·E·F·G·H·I·J·K·L·M·N·O·P·Q·R·S·T·U·V·W·X·Y·Z
A

E. abdita – E. adelbertensis – E. aipo – E. aipomek – E. akameku – E. alexanderi – E. allerbergeri – E. amabilis – E. ambua – E. andakombensis – E. anggiensis – E. apistefti – E. arfakensis – E. ascendens – E. aseki – E. astrophallus – E. athesphatos – E. atowaso – E. atra – E. atrata – E. atripennis – E. aubei – E. australiae – E. australis –
B

E. bacchusi – E. b. herzogensis – E. bagus – E. baliem – E. barbarae – E. bewani – E. bewaniensis – E. bifida – E. bimaculata – E. bismarckensis – E. boulevardi – E. brahminensis – E. brazza – E. broschii – E. brownei – E. brunoi – E. bundiensis – E. burwelli –
C

E. casuarina – E. charlottae – E. commatifera – E. craterensis – E. creuxorum – E. cyclops –
D

E. damantiensis – E. danae – E. desii –
E

E. edeltraudae – E. ekari – E. elongatula – E. eme – E. ekpliktiki – E. erteldi – E. evelyncheesmanae –
F

E. ferruginea – E. feryi – E. flammi – E. foja – E. fume –
G

E. gapa – E. garaina – E. gaulorum – E. gelima – E. glypta – E. gorokaensis – E. gracilis –
H

E. haia – E. hansferyi – E. heidiae – E. herowana – E. hintelmannae – E. hudsoni –
I

E. ibalimi – E. inexspectata – E. injiensis – E. interrupta – E. iratoi – E. irianensis –
J

E. jaseminae – E. jeannae – E. jimiensis –
K

E. kabwumensis – E. kailaki – E. kainantuensis – E. kakapupu – E. karmurensis – E. keki – E. ketembang – E. kinibeli – E. kisli – E. knoepfchen – E. kobau – E. koghis – E. kolleri – E. koroba – E. kowalskii – E. ksionseki – E. kumulensis –
L

E. larsoni – E. leae – E. lembena – E. likui – E. lilianae –
M

E. maculata – E. madangensis – E. manfredi – E. manokwariensis – E. mantembu – E. marawaga – E. marinae – E. may – E. me – E. mekilensis – E. melanaria – E. mendiensis – E. menyamya – E. messeri – E. mianminensis – E. michaelensis – E. mimika – E. miriae – E. monae – E. mondmillensis – E. monteithi – E. morobensis – E. munaso –
N

E. nehoue – E. nielsi – E. niklasi – E. nomax – E. novaecaledoniae –
O

E. oceai – E. oiwa – E. okapa – E. okbapensis – E. o. hajeki – E. oksibilensis – E. oraia – E. ouin –
P

E. parvula – E. patepensis – E. perfecta – E. pinocchio – E. piusi – E. poellabauerae – E. polita – E. posmani – E. pseudoastrophallus – E. pseudobifida – E. pseudoedeltraudae – E. pseudoeme – E. pseudofume – [[sp|E|xocelina|pseudojaseminae}} E. pseudomarinae – E. pseudopusilla – E. pseudosoppi – E. pui – E. pulchella – E. pulukensis – E. punctipennis – E. pusilla –
R

E. ransikiensis – E. rasilis – E. rasjadi – E. remyi – E. riberai – E. rivula – E. rotteri – E. rufa –
S

E. saltusholmesensis – E. sanctimontis – E. sandaunensis – E. schoelleri – E. shizong – E. sima – E. simbaiarea – E. simbaiensis – E. simbaijimi – E. simoni – E. simplex – E. skalei – E. soppi – E. staneki – E. subjecta – E. sugayai – E. sumokedi –
T

E. tabubilensis – E. takime – E. talaki – E. tariensis – E. tarmluensis – E. tekadu – E. tomhansi – E. tsinga –
U

E. ullrichi – E. unipo – E. utowaensis –
v

E. varirata – E. vladimiri – E. vovai –
W

E. waaf – E. waigeoensis – E. wannangensis – E. warahulenensis – E. warasera – E. wareaga – E. weylandensis – E. wigodukensis – E. woitapensis – E. wondiwoiensis –
Y

E. yoginofi
Name

Exocelina Broun, 1886
Original status: valid genus
Gender: feminine
Type species: Exocelina advena Broun, 1886 [=Celina australis Clark, 1863], by monotypy

Synonyms

Papuadytes Balke, 1998
Original status: valid subgenus of Copelatus
Gender: masculine
Type species: Copelatus rivulus Balke, 1998, by original designation
Synonymised by: Nilsson & Fery (2006, implied), Nilsson (2007)

References
Primary references

Balke, M. 1998: Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): the running water species, part 1. Annalen des Naturhistorischen Museums in Wien, 100B: 301–341. ISSN: 0255-0105 PDF

Additional references

Balke, M. & Ribera, I. 2020. A subterranean species of Exocelina diving beetle from the Malay Peninsula filling a 4,000 km distribution gap between Melanesia and southern China. Subterranean Biology, 34: 25–37. DOI: 10.3897/subtbiol.34.50148 Open access Reference page.
Broun, T. 1886: Manual of the New Zealand Coleoptera. Parts III and IV. Colonial Museum and Geological Survey Department, Wellington. Internet Archive BHL BUGZ
Clark, H. 1863: Catalogue of the Dytiscidae and Gyrinidae of Australasia with descriptions of new species. Journal of entomology, 2: 14–23. Internet Archive BHL
Nilsson, A.N. 2007: Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998. Latissimus, 23: 33–34. ISSN: 0966-2235
Nilsson, A.N.; Fery, H. 2006: World Catalogue of Dytiscidae – corrections and additions, 3 (Coleoptera: Dytiscidae). Koleopterologische rundschau, 76: 55–74. ISSN: 0075-6547
Shaverdo, H.V. et al. 2012: Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1-76. DOI: 10.3897/zookeys.250.3715 Reference page.
Shaverdo, H.V., Sagata, K., Panjaitan, R., Menufandu, H., & Balke, M. (2014). Description of 23 new species of the Exocelina ekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 468: 1-83. DOI: 10.3897/zookeys.468.8506 Reference page.
Shaverdo, H.V., Panjaitan, R. & Balke, M. 2016. A new, widely distributed species of the Exocelina ekari-group from West Papua (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 554: 69-85. DOI: 10.3897/zookeys.554.6065. Reference page.
Shaverdo, H.; Panjaitan, R.; Balke, M. 2016: Exocelina ransikiensis sp. nov. from the Bird’s Head of New Guinea (Coleoptera: Dytiscidae: Copelatinae). Acta Entomologica Musei Nationalis Pragae 56(1): 103-108. Full article (PDF) Reference page.
Shaverdo, H.V., Sagata, K. & Balke, M. 2016. Description of two new species of the Exocelina broschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 577: 125–148. DOI: 10.3897/zookeys.577.7254 Reference page.
Shaverdo, H.V., Sagata, K. & Balke, M. 2016. Taxonomic revision of New Guinea diving beetles of the Exocelina danae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 619: 45–102. DOI: 10.3897/zookeys.619.9951 Reference page.
Shaverdo, H., Sagata, K. & Balke, M. 2018. Introduction of the Exocelina casuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). Zookeys 803: 7–70. DOI: 10.3897/zookeys.803.28903 Reference page.
Shaverdo, H.V., Sumoked, B. & Balke, M. 2017. Descriptions of two new species and one new subspecies from the Exocelina okbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 715: 17—37. DOI: 10.3897/zookeys.715.15913. Reference page.
Shaverdo, H., Surbakti, Sumoked, B. & Balke, M. 2020. Two new species of the Exocelina ekari group from New Guinea with strongly modified male antennae (Coleoptera, Dytiscidae, Copelatinae). ZooKeys, 960: 63–78. DOI: 10.3897/zookeys.960.55007 Open access Reference page.
Shaverdo, H.V., Surbakti, S., Sumoked, B. & Balke, M. 2021. Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae). Zookeys 1026: 45–67. DOI: 10.3897/zookeys.1026.61554 Open access Reference page.
Shaverdo, H., Wild, M., Sumoked, B. & Balke, M. 2017. Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 665: 93—120. DOI: 10.3897/zookeys.665.11792. Reference page.
Shaverdo, H., Surbakti, S., Warikar, E.L., Sagata, K. & Balke, M. 2019. Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys, 878: 73–143. DOI: 10.3897/zookeys.878.37403 Open access Reference page.
Watts, C.H.S., Hendrich, L. & Balke, M. 2016. A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae). Subterranean Biology, 19: 23–29. DOI: 10.3897/subtbiol.19.9513 Open access Reference page.
Watts, C.H.S.; Humphreys, W.F. 2009: Fourteen new Dytiscidae (Coleoptera) of the genera Limbodessus Guignot, Paroster Sharp, and Exocelina Broun from underground waters in Australia. Transactions of the Royal Society of South Australia, 133: 62–107. ISSN: 0372-1426
Wewalka, G.; Balke, M.; Hendrich, L. 2010: Dytiscidae: Copelatinae (Coleoptera). Pp. 45-128 in: Jäch, M.A. & Balke, M. (eds.): Water beetles of New Caledonia (part 1). Monographs on Coleoptera, 3 . ISSN: 1027-8869

Additional references

Balke, M. 1999: Two new species of the genus Copelatus Erichson, 1832, subgenus Papuadytes Balke, 1998 (Insecta: Coleoptera: Dytiscidae) from Papua New Guinea. Annalen des Naturhistorischen Museums in Wien, 101B: 273–276. ISSN: 0255-0105 PDF
Balke, M. 2001: Replacement names for three New Guinea species of Copelatus, subgenus Papuadytes Balke, 1998 (Coleoptera: Dytiscidae). Annalen des Naturhistorischen Museums in Wien, 103B: 361–362. ISSN: 0255-0105 PDF
Balke, M.; Bergsten, J. 2003: Dytiscidae: I. Copelatus (Papuadytes) shizong sp. n. from Yunnan (China), the first member of Papuadytes Balke found west of the Wallace Line (Coleoptera). Pp. 89-94 in Jäch, M.A.; Ji, L. (eds.) Water beetles of China. Volume 3. Zoologisch - Botanische Gesellschaft, Vienna. [not seen]
Balke, M.; Pons, J.; Ribera, I.; Sagata, K.; Vogler, A.P. 2007: Infrequent and unidirectional colonization of hyperdiverse Papuadytes diving beetles in New Caledonia and New Guinea. Molecular phylogenetics and evolution, 42: 505–516. DOI: 10.1016/j.ympev.2006.07.019 PDF
Balke, M.; Ribera, I.; Vogler, A.P. 2004: MtDNA phylogeny and biogeography of Copelatinae, a highly diverse group of tropical diving beetles (Dytiscidae). Molecular phylogenetics and evolution, 32: 866–880. DOI: 10.1016/j.ympev.2004.03.014
Balke, M.; Watts, C.H.S.; Cooper, S.J.B.; Humphreys, W.F.; Vogler, A.P. 2004: A highly modified stygobiont diving beetle of the genus Copelatus (Coleoptera, Dytiscidae): taxonomy and cladistic analysis based on mitochondrial DNA sequences. Systematic entomology, 29: 59–67. DOI: 10.1111/j.1365-3113.2004.00229.x
Fikáček, M.; Hájek, J.; Schmied, H. 2011: On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae). ZooKeys, 78: 15–25. ISSN: 1313-2970 (online) ISSN: 1313-2989 (print) DOI: 10.3897/zookeys.78.800
Nilsson, A.N. 2001: Dytiscidae (Coleoptera). World catalogue of insects, 3 . Stenstrup: Apollo Books. ISBN 8788757625 [not seen]
Shaverdo, H.V.; Sagata, K.; Balke, M. 2005: Five new species of the genus Papuadytes Balke, 1998 from New Guinea (Coleoptera: Dytiscidae). Aquatic insects, 27: 269–280. DOI: 10.1080/01650420500290169
Shaverdo, H., Surbakti, S., Sumoked, B. & Balke, M. 2020. Three new species of Exocelina Broun, 1886 from the southern slopes of the New Guinea central range, with introduction of the Exocelina skalei group (Coleoptera, Dytiscidae, Copelatinae. ZooKeys, 1007: 129–143. DOI: 10.3897/zookeys.1007.59351 Open access Reference page.

Exocelina is a genus of beetles in the family Dytiscidae, containing the following species:

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler, 2004)
Exocelina aipo (Balke, 1998)
Exocelina aipomek (Balke, 1998)
Exocelina alexanderi Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina ascendens (Balke, 1998)
Exocelina astrophallus (Balke, 1998)
Exocelina atowaso Shaverdo, Sagata & Balke, 2005
Exocelina atra (Sharp, 1882)
Exocelina aubei (Montrouzier, 1860)
Exocelina atratus (J.Balfour-Browne, 1939)
Exocelina atripennis (J.Balfour-Browne, 1939)
Exocelina australiae (Clark, 1863)
Exocelina australis (Clark, 1863)
Exocelina bacchusi (Balke, 1998)
Exocelina bagus (Balke & Hendrich, 2001)
Exocelina baliem (Hendrich & Balke, 2013)
Exocelina bewaniensis Shaverdo, Menufandu & Balke, 2014
Exocelina bifida Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina bimaculata (Perroud & Montrouzier, 1864)
Exocelina bismarckensis Shaverdo & Balke, 2014
Exocelina boulevardi (Watts, 1978)
Exocelina brahminensis Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina broschii (Balke, 1998)
Exocelina brownei (Guignot, 1942)
Exocelina brunoi Wewalka, Balke & Hendrich, 2010
Exocelina casuarina (Balke & Hendrich, 1998)
Exocelina commatifera (Heller, 1916)
Exocelina craterensis Shaverdo & Balke, 2014
Exocelina creuxorum Wewalka, Balke & Hendrich, 2010
Exocelina damantiensis (Balke, 1998)
Exocelina danae (Balke, 1998)
Exocelina elongatula (W.J. Macleay, 1871)
Exocelina erteldi (Balke, 1998)
Exocelina evelyncheesmanae Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina ferruginea (Sharp, 1882)
Exocelina feryi Wewalka, Balke & Hendrich, 2010
Exocelina flammi Wewalka, Balke & Hendrich, 2010
Exocelina gapa (Watts, 1978)
Exocelina gaulorum Wewalka, Balke & Hendrich, 2010
Exocelina gelima Wewalka, Balke & Hendrich, 2010
Exocelina glypta (Guignot, 1955)
Exocelina gorokaensis Shaverdo & Balke, 2014
Exocelina gracilis (Sharp, 1882)
Exocelina hansferyi Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina heidiae (Balke, 1998)
Exocelina herowana Shaverdo & Balke, 2014
Exocelina hintelmannae Shaverdo, Sagata & Balke, 2005
Exocelina inexspectata Wewalka, Balke & Hendrich, 2010
Exocelina interrupta (Perroud & Montrouzier, 1864)
Exocelina irianensis Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina jaseminae (Balke, 1998)
Exocelina jimiensis Shaverdo & Balke, 2014
Exocelina kainantuensis (Balke, 2001)
Exocelina kakapupu Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina karmurensis (Balke, 1998)
Exocelina ketembang (Balke, 1998)
Exocelina kinibeli Shaverdo & Balke, 2014
Exocelina kisli Shaverdo & Balke, 2014
Exocelina knoepfchen Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina koghis Wewalka, Balke & Hendrich, 2010
Exocelina kolleri Wewalka, Balke & Hendrich, 2010
Exocelina ksionseki Shaverdo & Balke, 2014
Exocelina larsoni (Balke, 1998)
Exocelina leae Wewalka, Balke & Hendrich, 2010
Exocelina lembena Shaverdo & Balke, 2014
Exocelina lilianae Wewalka, Balke & Hendrich, 2010
Exocelina maculata (Sharp, 1882)
Exocelina madangensis (Balke, 2001)
Exocelina manfredi (Balke, 1998)
Exocelina mantembu Shaverdo & Balke, 2014
Exocelina me (Balke, 1998)
Exocelina melanaria (Sharp, 1882)
Exocelina messeri (Balke, 1999)
Exocelina michaelensis Shaverdo & Balke, 2014
Exocelina miriae (Balke, 1998)
Exocelina monae (Balke, 1998)
Exocelina monteithi Wewalka, Balke & Hendrich, 2010
Exocelina munaso Shaverdo, Sagata & Balke, 2005
Exocelina nielsi Wewalka, Balke & Hendrich, 2010
Exocelina niklasi Wewalka, Balke & Hendrich, 2010
Exocelina nomax (J.Balfour-Browne, 1939)
Exocelina novaecaledoniae (J.Balfour-Browne, 1939)
Exocelina oceai Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina ouin Wewalka, Balke & Hendrich, 2010
Exocelina parvulus (Boisduval, 1835)
Exocelina patepensis (Balke, 1998)
Exocelina perfectus (Sharp, 1882)
Exocelina pinocchio Shaverdo & Balke, 2014
Exocelina poellabauerae Wewalka, Balke & Hendrich, 2010
Exocelina polita (Sharp, 1882)
Exocelina pseudoastrophallus Shaverdo & Balke, 2014
Exocelina pseudobifida Shaverdo & Balke, 2014
Exocelina pseudoedeltraudae Shaverdo & Balke, 2014
Exocelina pseudosoppi Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina punctipennis (Lea, 1899)
Exocelina rasjadi Watts & Humphreys, 2009
Exocelina remyi Wewalka, Balke & Hendrich, 2010
Exocelina rasilis (Lea, 1899)
Exocelina rivula (Balke, 1998)
Exocelina rotteri Wewalka, Balke & Hendrich, 2010
Exocelina rufa (Balke, 1998)
Exocelina sanctimontis (Balke, 1998)
Exocelina schoelleri Wewalka, Balke & Hendrich, 2010
Exocelina shizong (Balke & Bergsten, 2003)
Exocelina simbaiarea Shaverdo & Balke, 2014
Exocelina simoni Wewalka, Balke & Hendrich, 2010
Exocelina simplex (Clark, 1863)
Exocelina skalei Shaverdo & Balke, 2014
Exocelina soppi Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina staneki Wewalka, Balke & Hendrich, 2010
Exocelina subjecta (Sharp, 1882)
Exocelina takime (Balke, 1998)
Exocelina talaki (Balke, 1998)
Exocelina tariensis Shaverdo & Balke, 2014
Exocelina tarmluensis (Balke, 1998)
Exocelina ullrichi (Balke, 1998)
Exocelina unipo Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina vladimiri Shaverdo, Sagata & Balke, 2005
Exocelina vovai Shaverdo & Balke, 2014
Exocelina waigeoensis Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina wannangensis Shaverdo & Balke, 2014
Exocelina weylandensis Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012
Exocelina wondiwoiensis Shaverdo, Hendrich & Balke in Shaverdo, Surbakti, Hendrich & Balke, 2012

References