Cossinae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Superfamilia: Cossoidea

Familia: Cossidae
Subfamilia: Cossinae
Tribus: Groenendaelini - Holcocerini - Zeuzerocossini
Overview of genera

Acossus – Afrikanetz – Afroarabiella – Aholcocerus – Alcterogystia – Arctiocossus – Assegaj – Bifiduncus – Brachygystia – Brachylia – Butaya – Camellocossus – Chinocossus – Citharalia – Comadia – Coryphodema – Cossodes – Cossus – Culama – Danielostygia – Dervishiya – Dieida – Dyspessa – Dyspessacossus – Endagria – Eogystia – Eremocossus – Fania – Garuda – Gobibatyr – Groenendaelia – Gumilevia – Hirtocossus – Hollowiella – Isoceras – Isocossus – Koboldocossus – Kotchevnik – Lamellocossus – Lichtensteiniana – Macrocossus – Macrocyttara – Mahommedella – Miacora – Mirocossus – Mormogystia – Namibiocossus – Nirvana (Yakovlev) – Paracossus – Paracossulus – Parahypopta – Patopta – Patoptoformis – Pecticossus – Planctogystia – Prionoxystus – Psychidocossus – Pygmeocossus – Rambuasalama – Rasa – Rethona – Roepkiella – Ronaldocossus – Samagystia – Semitocossus – Stroehleia – Stygioides – Sundacossus – Toronia (Cossidae) – Vartiania – Wiltshirocossus – Wittocossus – Zeuzerocossus – Zyganisus
Name

Cossinae Leach, [1815] 1830

Type genus: Cossus Fabricius, 1793
References

Daniel, F., 1971: Österreichische Expeditionen nach Persien und Afghanistan. Beiträge zur Lepidopterenfauna, Teil 16 (3. Beitrag zur Bombyces- und Sphinges-Fauna). Annalen des Naturhistorische Museum in Wien 75: 651–660. Full article: [1].
Kallies, A. & D.J. Hilton, 2012: Revision of Cossinae and small Zeuzerinae from Australia (Lepidoptera: Cossidae). Zootaxa 3454: 1–62. Abstract: [2].
Schoorl, J.W., 1990: A phylogenetic study on Cossidae (Lepdoptera: Ditrysia) based on external adult morphology. Zoologische Verhandelingen 263: 1–295. Full article: [3].
Yakovlev, R.V., 2004: Cossidae of Thailand. Part 1. (Lepidoptera: Cossidae). Atalanta 35 (3-4): 335–351.
Yakovlev, R.V. 2004: Two new genera of Carpenter Moths from the Palaearctic (Lepidoptera: Cossidae). Atalanta 35 (3/4): 357–368.
Yakovlev, R.V., 2004: Cossidae of Thailand. Part 2. (Lepidoptera: Cossidae). Atalanta 35 (3-4): 383–389.
Yakovlev, R. V., 2005: Cossidae (Lepidoptera) of Andaman Islands (India). Tinea 18 (4): 257–260.
Yakovlev, R. V., 2006: New Cossidae (Lepidoptera) from Asia, Africa and Macronesia, Tinea 19 (3): 188–213.
Yakovlev, R.V., 2009: New taxa of African and Asian Cossidae (Lepidoptera). Euroasian Entomological Journal 8 (3): 353–361. Full article: [4].
Yakovlev, R.V., 2011: Catalogue of the Family Cossidae of the Old World. Neue Entomologische Nachrichten, 66: 1–129.
Yakovlev, R.V. 2021f. New genus of Cossinae (Lepidoptera: Cossidae) from the Republic of South Africa. Russian Entomological journal 30(2): 175–177. DOI: 10.15298/rusentj.30.2.11. Open access. Reference page.
Yakovlev, R.V. & A. Saldaitis, 2008: Rambuasalama augustasi gen. nov. and spec. nov. of Cossidae from Madagascar (Lepidoptera: Cossidae). Atalanta 39 (1-4): 401–403.
Yakovlev, R.V. & T.J. Witt, 2009: The Carpenter Moths (Lepidoptera:Cossidae) of Vietnam. Entomofauna Supplement 16: 11–32.
Yakovlev, R.V., Pljustch, I.G., Skrylnik, Y., Pak, O.V. & Witt, T.J. 2015. The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park. Zootaxa 3990(1): 41–72. DOI: 10.11646/zootaxa.3990.1.3. Preview (PDF) Reference page.

The Cossinae are the nominate subfamily of the Cossidae (carpenter or goat moths). The caterpillars of several Cossinae species, such as the carpenterworm (Prionoxystus robiniae) and the goat moth (Cossus cossus), are significant pests. On the other hand, in Chile the caterpillars of the Chilean moth (Chilecomadia moorei) are collected on a commercial scale for sale as fishing bait and terrarium pet food; they are usually called "butterworms" in international trade.

The Cossulinae have been separated from the Cossinae in recent decades, but this was not universally accepted at first. Some misplaced genera have been moved between the subfamilies, and as it seems the Cossulinae at least now represent a monophyletic group.
Contents

1 Systematics
2 Former genera
3 Footnotes
4 References

Systematics

Some significant species are also listed:[1]

Unplaced to tribe/placement unknown
Citharalia Clench, 1957 (formerly in Cossulinae)
Comadia Barnes & McDunnough, 1911
Cossodes White, 1841
Culama Walker, 1856
Dieidapersa Strand in Stichel, 1911
Dieida Strand, 1911
Fania Barnes & McDunnough, 1911 (tentatively placed here)
Idioses Turner, 1927
Macrocyttara Turner, 1918
Miacora Dyar, 1905 (tentatively placed here)
Prionoxystus Grote, 1882
Psychidocossus D. S. Fletcher, 1982
Surcossus Heimlich, 1960
Trigena Dyar, 1905 (formerly in Cossulinae)
Zyganisus Viette, 1951
Zeuzerocossini Yakovlev, 2008
Acossus Dyar, 1905
Alcterogystia Schoorl, 1990
Arctiocossus Felder, 1874
Assegaj Yakovlev, 2006
Brachygystia Schoorl, 1990
Chingizid Yakovlev, 2011
Chinocossus Yakovlev, 2006
Dervishiya Yakovlev, 2006
Eogystia Schoorl, 1990
Eremocossus Hampson, 1893
Gobibatyr Yakovlev, 2004
Kalimantanossus Yakovlev, 2011
Mahommedella Yakovlev, 2011
Neurocossus Yakovlev, 2010
Paropta Staudinger, 1900
Patoptoformis Yakovlev, 2006
Planctogystia Schoorl, 1990
Rethona Walker, 1855
Ronaldocossus Yakovlev, 2006
Wiltshirocossus Yakovlev, 2007
Zeuzerocossus Yakovlev, 2008
Endagriini Duponchel, [1845]
Afroarabiella Yakovlev, 2008
Dyspessa Hübner, 1820
Groenendaelia Yakovlev, 2004
Hollowiella Yakovlev & Witt, 2009
Isoceras Turati, 1924
Paracossulus Schoorl, 1990
Stygoides Bruand, 1853
Samagystia Schoorl, 1990
Holcocerini Yakovlev, 2006
Afrikanetz Yakovlev, 2009
Aholcocerus Yakovlev, 2006
Barchaniella Yakovlev, 2006
Brachylia Felder, 1874
Camellocossus Yakovlev, 2011
Coryphodema Felder, 1874
Cossulus Staudinger, 1887
Cossus Fabricius, 1794
Cryptoholcocerus Yakovlev, 2006
Deserticossus Yakovlev, 2006
Dyspessacossus Daniel, 1953
Franzdanielia Yakovlev, 2006
Gumilevia Yakovlev, 2011
Hirtocossus Schoorl, 1990
Holcocerus Staudinger, 1884
Isocossus Roepke, 1957
Kerzhnerocossus Yakovlev, 2011
Koboldocossus Yakovlev, 2011
Kotchevnik Yakovlev, 2004
Mirocossus Schoorl, 1990
Macrocossus Aurivillius, 1900
Mormogystia Schoorl, 1990
Paracossus Hampson, 1904
Parahypopta Daniel, 1961
Plyustchiella Yakovlev, 2006
Pygmeocossus Yakovlev, 2005
Rambuasalama Yakovlev & Saldaitis, 2008
Reticulocossus Yakovlev, 2011
Roepkiella Yakovlev & Witt, 2009
Semitocossus Yakovlev, 2007
Streltzoviella Yakovlev, 2006
Sundacossus Yakovlev, 2006
Vartiania Yakovlev, 2004
Wittocossus Yakovlev, 2004
Yakudza Yakovlev, 2006

Former genera

Bifiduncus
Butaya Yakovlev, 2004
Catopta Staudinger, 1899
Chiangmaiana Kemal & Koçak, 2005 (≡Nirvana Yakovlev, 2004 nom. invalid.(preoc.name) non Stål, 1859 nec Kirkaldy, 1900 nec Yakovlev, 2007 nec Tsukuda & Nishiyama, 1979);(≡Nirrvanna Yakovlev, 2007 nom. invalid.( junior object. syn.), objective replacement name (mistaken repl.n.))
Chilecomadia
Danielostygia Reisser, 1962 (tentatively placed here)
Lamellocossus Daniel, 1956 (tentatively placed here)
Toronia Barnes & McDunnough, 1911
Yakovlevina Kemal & Koçak, 2005 (=Garuda Yakovlev, 2005)
Stygia Latreille, 1803 (tentatively placed here)

Footnotes

See references in Savela (2008)

References

Savela, Markku (2008): Lepidoptera and Some Other Life Forms Cossinae. Version of 17 July 2008. Retrieved 27 February 2009.
Ревизия древоточцев рода Holcocerus Staudinger, 1884 (s. l.)