Copelatus

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Adephaga

Familia: Dytiscidae
Subfamilia: Copelatinae
Genus: Copelatus
Species: C. amphibius – C. ankaratra – C. babyrousa – C. baranensis – C. betampona – C. bezdeki – C. biroi – C. bougainvillensis – C. boukali – C. brendelli – C. cessaima – C. chevrolati – C. commatifer – C. davidi – C. deccanensis – C. djiboutensis – C. doriae – C. gestroi – C. haemorrhoidalis – C. glyphicus – C. imasakai – C. kaszabi – C. kely – C. kietensis – C. laevipennis – C. luteomaculatus – C. marginatus – C. martinbaehri – C. martini – C. masculinus – C. maushomi – C. portitor – C. posticatus – C. pseudostriatus – C. safiotra – C. sibelaemontis – C. strigosulus – C. urceolus – C. variistriatus – C. vokoka – C. wayanadensis – C. weymarni – †C. aphroditae – †C. convexus – †C. fossilis – †C. ponomarenkoi – †C. predaveterus – †C. stavropolitanus – C. vancli – C. zanatanensis
...
Name

Copelatus Erichson, 1832
References

Erichson W.F., 1832. Genera dyticeorum.
Fikáček, M.; Hájek, J.; Schmied, H. 2011: On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae). ZooKeys, 78: 15–25. DOI: 10.3897/zookeys.78.800
Hájek, J., Hendrich, L. & Balke, M. 2018. The Copelatus doriae-masculinus species complex in Greater Sunda, with description of three new species and a new synonymy' Spixiana 41(1): 77-90. Full article (PDF). Reference page.
Hájek, J., Shaverdo, H.V., Hendrich, L. & Balke, M. 2021. A review of Copelatus diving beetles from the Solomon Islands, reporting the discovery of six new species (Coleoptera, Dytiscidae, Copelatinae). Zookeys 1023: 81–118. DOI: 10.3897/zookeys.1023.61478 Open access Reference page.
Hendrich, L., Shaverdo, H., Hájek, J. & Balke, M. 2019. Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys, 889: 81–152. DOI: 10.3897/zookeys.889.39090 Open access Reference page.
Larson D.J., Alarie Y., Roughley R.E. 2000: Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska, National research Council of Canada, Ottawa.
Manuel, M., Deler-Hernández, A., Megna, Y.S. & Hájek, J. 2018. Copelatus Erichson from the Dominican Republic, with the description of a new species, comments on elytral striation and faunistic notes on Antillean species (Coleoptera: Dytiscidae: Copelatinae). Zootaxa 4399(3): 371–385. DOI: 10.11646/zootaxa.4399.3.7 Reference page.
Megna, Y.S.; Epler, J.H. 2012: A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae). Acta Entomologica Musei Nationalis Pragae 52(2): 383–410. PDF Reference page.
Michat, M.C.; Torres, P.L.M. 2009: A preliminary study on the phylogenetic relationships of Copelatus Erichson (Coleoptera: Dytiscidae: Copelatinae) based on larval chaetotaxy and morphology. Hydrobiologia, 632: 309–327. DOI: 10.1007/s10750-009-9853-2
Miller, K.B; Balke, M. 2003: The unusual occurrence of aquatic beetles in amber, Copelatus aphroditae Balke, n. sp. and C. predaveterus Miller, n. sp., (Coleoptera: Dytiscidae: Copelatinae). Proceedings of the Entomological Society of Washington, 105(4): 809–815. BHL Internet Archive PDF
Ranarilalatiana, T. & Bergsten, J. 2019. Discovery of a specialist Copelatinae fauna on Madagascar: highly ephemeral tropical forest floor depressions as an overlooked habitat for diving beetles (Coleoptera, Dytiscidae). Zookeys, 871: 89–118. DOI: 10.3897/zookeys.871.36337 Reference page.
Ranarilalatiana, T., Ravaomanarivo, L.H.R. & Bergsten, J. 2019. Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species Zookeys, 869: 19–90. DOI: 10.3897/zookeys.869.33997 Reference page.
Shaverdo, H.; Monaghan, M.T.; Lees, D.C.; Ranaivosolo, R.; Balke, M. 2008: Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae). Systematics and biodiversity, 6: 43–51. DOI: 10.1017/S1477200007002599
Sheth, S.D., Ghate, H.V. & Hájek, J. 2018. Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). Zootaxa 4459(2): 235–260. DOI: 10.11646/zootaxa.4459.2.2 Paywall Reference page.
Wewalka, G.; Balke, M.; Hendrich, L. 2010: Dytiscidae: Copelatinae (Coleoptera). Pp. 45-128 in: Jäch, M.A. & Balke, M. (eds.): Water beetles of New Caledonia (part 1). Monographs on Coleoptera, 3. ISSN: 1027-8869

Links

ION
Nomenclator Zoologicus
Tribe Copelatini (atlas of beetles of Russia) [1]

Copelatus is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia.
Contents

1 Systematics
2 Species
3 References
4 External links

Systematics

The genus Copelatus is divided into several subgenera (Colepatus, Papuadytes etc.), some of which are sometimes treated as separate genera. It may be paraphyletic with respect to the smaller Copelatinae genera Lacconectus and Aglymbus.[2] The species include:[3]
Species

Copelatus abonnenci Guignot, 1939
Copelatus acamas Guignot, 1955
Copelatus advena Sharp, 1882
Copelatus aemulus Bilardo & Rocchi, 1995
Copelatus aequatorius Régimbart, 1899
Copelatus aethiopicus Régimbart, 1906
Copelatus agrias Guignot, 1954
Copelatus aldabricus J.Balfour-Browne, 1950
Copelatus alternatus Sharp, 1882
Copelatus amaroides Guignot, 1952
Copelatus amatolensis Omer-Cooper, 1965
Copelatus amazonicus Régimbart, 1889
Copelatus ambiguus Bertrand & Legros, 1975
Copelatus anastomosans Guignot, 1952
Copelatus andobonicus Guignot, 1960
Copelatus andreinii Régimbart, 1905
Copelatus andrewesi J.Balfour-Browne, 1939
Copelatus angolensis Peschet, 1924
Copelatus angustus Gschwendtner, 1932
Copelatus annobomensis J.Balfour-Browne, 1939
Copelatus anthracinus Régimbart, 1895
Copelatus aphroditae Balke, 2003
Copelatus apicalis Fairmaire, 1898
Copelatus apuzzoi Bilardo & Rocchi, 1999
Copelatus aruensis J.Balfour-Browne, 1939
Copelatus assamensis Vazirani, 1970
Copelatus assimilis Régimbart, 1895
Copelatus ateles Guignot, 1955
Copelatus atrosulcatus Régimbart, 1906
Copelatus bacchusi Wewalka, 1981
Copelatus bacillifer Guignot, 1955
Copelatus baculiformis Guignot, 1955
Copelatus badeni Sharp, 1882
Copelatus bakewelli J.Balfour-Browne, 1939
Copelatus bangalorensis Vazirani, 1970
Copelatus baoulicus Bilardo & Pederzani, 1978
Copelatus barbouri Young, 1942
Copelatus basifasciatus Régimbart, 1895
Copelatus basilewskyi Bilardo & Pederzani, 1979
Copelatus bechynei Guignot, 1953
Copelatus befasicus Guignot, 1956
Copelatus bengalensis Guignot, 1955
Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996
Copelatus bibulus Guignot, 1948
Copelatus biformis Sharp, 1882
Copelatus bilunatus Guignot, 1955
Copelatus binaghii Bilardo & Pederzani, 1978
Copelatus biolleyi Guignot, 1952
Copelatus biroi Guignot, 1956
Copelatus biseriatus J.Balfour-Browne, 1939
Copelatus biswasi Mukherjee & Sengupta, 1986
Copelatus blancasi Guignot, 1958
Copelatus blatchleyi Young, 1953
Copelatus bolivianus Guignot, 1957
Copelatus bombycinus Guignot, 1952
Copelatus bonvouloiri Sharp, 1882
Copelatus bottegoi Régimbart, 1895
Copelatus boukali Hendrich & Balke, 1998
Copelatus brancuccii Rocchi, 1979
Copelatus brasiliensis Zimmermann, 1921
Copelatus brevistrigatus Guignot, 1959
Copelatus brivioi Rocchi, 1976
Copelatus brullei Aubé, 1838
Copelatus brunneus J.Balfour-Browne, 1939
Copelatus buqueti Aubé, 1838
Copelatus burgeoni Gschwendtner, 1930
Copelatus caelatipennis Aubé, 1838
Copelatus caelatus Guignot, 1952
Copelatus caffer J.Balfour-Browne, 1939
Copelatus camerunensis Guignot, 1941
Copelatus capensis Sharp, 1882
Copelatus carayoni Legros, 1949
Copelatus carinatus Sharp, 1882
Copelatus celinoides Guignot, 1952
Copelatus cessaima Caetano, de C. Bena, & Vanin, 2013[4]
Copelatus ceylonicus Vazirani, 1969
Copelatus cheesmanae J.Balfour-Browne, 1939
Copelatus chevrolati Aubé, 1838
Copelatus chibcha Guignot, 1952
Copelatus chipiriricus Guignot, 1957
Copelatus chloroticus Régimbart, 1899
Copelatus cinnamomeus Régimbart, 1895
Copelatus clarki Sharp, 1882
Copelatus collarti Gschwendtner, 1932
Copelatus compertus Guignot, 1956
Copelatus concii Bilardo, 1982
Copelatus concolor Sharp, 1882
Copelatus concolorans J.Balfour-Browne, 1939
Copelatus confinis Bilardo & Rocchi, 1999
Copelatus congo Gschwendtner, 1938
Copelatus consimilis Bilardo & Rocchi, 2002
Copelatus consors Sharp, 1882
Copelatus cooperae Bilardo & Pederzani, 1972
Copelatus cordovai Megna & Epler, 2012[5]
Copelatus cordylinoides J.Balfour-Browne, 1938
Copelatus coxalis Sharp, 1882
Copelatus crassus Régimbart, 1895
Copelatus cryptarchoides Régimbart, 1899
Copelatus cubaensis Schaeffer, 1908
Copelatus curtistriatus Bilardo & Rocchi, 1995
Copelatus daemeli Sharp, 1882
Copelatus danyi Megna & Epler, 2012[6]
Copelatus darlingtoni Young, 1942
Copelatus debilis Sharp, 1882
Copelatus decellei Bilardo, 1982
Copelatus decemsulcatus Régimbart, 1895
Copelatus deceptor Bilardo & Rocchi, 1995
Copelatus depressus Sharp, 1882
Copelatus descarpentriesi Bertrand & Legros, 1975
Copelatus diffisus Guignot, 1939
Copelatus dimorphus Sharp, 1882
Copelatus distinctus Aubé, 1838
Copelatus distinguendus Régimbart, 1903
Copelatus divisus Watts, 1978
Copelatus dolosus Guignot, 1956
Copelatus doriae Sharp, 1882
Copelatus duodecimstriatus Aubé, 1838
Copelatus duponti Aubé, 1838
Copelatus edax Guignot, 1955
Copelatus efoutensis Bilardo & Rocchi, 1995
Copelatus ejactus Omer-Cooper, 1965
Copelatus ellai Bilardo & Rocchi, 1995
Copelatus elutus Guignot, 1958
Copelatus enganensis Guignot, 1940
Copelatus epactus Guignot, 1948
Copelatus erichsonii Guérin-Méneville, 1849
Copelatus esteriensis Bilardo & Pederzani, 1978
Copelatus eucritus Guignot, 1952
Copelatus evanidus Bilardo & Rocchi, 1995
Copelatus exaratus Sharp, 1882
Copelatus externus Kirsch, 1873
Copelatus fasciatus Bilardo & Rocchi, 1995
Copelatus fastidiosus Guignot, 1959
Copelatus feae Régimbart, 1888
Copelatus fernandensis Guignot, 1952
Copelatus ferruginicollis Régimbart, 1895
Copelatus ferus Guignot, 1953
Copelatus festae Régimbart, 1899
Copelatus fidschiensis Zimmermann, 1928
Copelatus fijiensis Guignot, 1955
Copelatus filiformis Sharp, 1882
Copelatus flavicans Guignot, 1952
Copelatus flavidus Régimbart, 1895
Copelatus fluviaticus Guignot, 1957
Copelatus fontanus J.Balfour-Browne, 1950
Copelatus fossilis Riha, 1974
Copelatus fractistriatus Bilardo & Rocchi, 1995
Copelatus freudei Guignot, 1955
Copelatus fryi J.Balfour-Browne, 1939
Copelatus fulviceps J.Balfour-Browne, 1938
Copelatus fuscipennis Sharp, 1882
Copelatus fuscomaculatus Guignot, 1952
Copelatus gabonicus Bilardo & Pederzani, 1978
Copelatus galapagoensis G.R.Waterhouse, 1845
Copelatus garambanus Guignot, 1955
Copelatus gardineri Scott, 1912
Copelatus geayi Régimbart, 1904
Copelatus geniculatus Sharp, 1882
Copelatus gentilis Sharp, 1882
Copelatus gestroi Régimbart, 1892
Copelatus gibsoni Vazirani, 1974
Copelatus glyphicus (Say, 1823)
Copelatus griffinii Régimbart, 1899
Copelatus gschwendtneri Guignot, 1939
Copelatus guadelupensis Legros, 1948
Copelatus guerini Aubé, 1838
Copelatus guineensis Guignot, 1955
Copelatus hararensis Guignot, 1952
Copelatus hardenbergi J.Balfour-Browne, 1950
Copelatus hebeter Guignot, 1953
Copelatus heterogynus Régimbart, 1899
Copelatus hydroporoides (Murray, 1859)
Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000
Copelatus iguelaensis Bilardo & Rocchi, 2002
Copelatus ilybioides Régimbart, 1895
Copelatus imasakai Matsui & Kitayama, 2000
Copelatus imitator Bilardo & Rocchi, 2002
Copelatus inaequalis Sharp, 1882
Copelatus incognitus Sharp, 1882
Copelatus indicus Sharp, 1882
Copelatus inopinatus Bilardo & Rocchi, 1995
Copelatus inornatus Sharp, 1882
Copelatus insidiosus Bilardo & Rocchi, 1995
Copelatus insolitus Chevrolat, 1863
Copelatus instabilis Régimbart, 1897
Copelatus instriatus Zimmermann, 1921
Copelatus insuetus Guignot, 1941
Copelatus insulanus Guignot, 1939
Copelatus integer Sharp, 1882
Copelatus internus Régimbart, 1904
Copelatus interstriatus J.Balfour-Browne, 1939
Copelatus inuber Guignot, 1952
Copelatus ipiformis Régimbart, 1895
Copelatus irinus Régimbart, 1899
Copelatus irregularis W.J. Macleay, 1871
Copelatus ischius Guignot, 1956
Copelatus jactator Guignot, 1955
Copelatus jamaicensis Young, 1942
Copelatus japonicus Sharp, 1884
Copelatus jarrigei Legros, 1954
Copelatus javanus Régimbart, 1883
Copelatus jocosus Guignot, 1955
Copelatus johannis Guignot, 1954
Copelatus kalaharii Gschwendtner, 1935
Copelatus kammuriensis Tamu & Tsukamoto, 1955
Copelatus karnatakus Holmen & Vazirani, 1990
Copelatus kaszabi Guignot, 1956
Copelatus kilimandjarensis Bilardo & Pederzani, 1972
Copelatus kindianus Guignot, 1954
Copelatus kongouensis Bilardo & Rocchi, 1999
Copelatus koreanus Mori, 1932
Copelatus laccophilinus Sharp, 1882
Copelatus laeticulus Sharp, 1882
Copelatus lamottei Legros, 1954
Copelatus lanzai Bilardo & Rocchi, 1995
Copelatus laraensis Bilardo & Rocchi, 1995
Copelatus lasckonyi Bilardo & Rocchi, 1995
Copelatus latens Guignot, 1952
Copelatus laticollis Régimbart, 1899
Copelatus latifasciatus Bilardo & Rocchi, 1999
Copelatus latipes Sharp, 1882
Copelatus latus J.Balfour-Browne, 1950
Copelatus leonardii Bilardo & Rocchi, 1999
Copelatus leonensis Legros, 1954
Copelatus lepersonneae Gschwendtner, 1943
Copelatus lignosus Guignot, 1955
Copelatus lineatipennis Guignot, 1955
Copelatus lineatus (Guérin-Méneville, 1838)
Copelatus longicornis Sharp, 1882
Copelatus lootensi Guignot, 1955
Copelatus louayensis Bilardo & Rocchi, 2004
Copelatus luctuosus Guignot, 1939
Copelatus luridescens Régimbart, 1889
Copelatus luteocinctus Guignot, 1955
Copelatus luteomaculatus Guignot, 1956
Copelatus luzonicus Guignot, 1952
Copelatus macellus Guignot, 1950
Copelatus madoni Guignot, 1952
Copelatus mahajanga Pederzani & Hájek, 2005
Copelatus mahleri Holmen & Vazirani, 1990
Copelatus makokouensis Bilardo & Rocchi, 1995
Copelatus malaisei Guignot, 1954
Copelatus mancus Sharp, 1887
Copelatus marginatus Sharp, 1882
Copelatus masculinus Régimbart, 1899
Copelatus massaicus Guignot, 1941
Copelatus mathani Guignot, 1952
Copelatus mbokoensis Bilardo & Rocchi, 2006
Copelatus melanogrammus Régimbart, 1883
Copelatus mimetes Guignot, 1957
Copelatus minimus Bilardo & Rocchi, 1995
Copelatus minor Bilardo & Pederzani, 1978
Copelatus minutissimus J.Balfour-Browne, 1939
Copelatus mocquerysi Régimbart, 1895
Copelatus mohelicus Guignot, 1959
Copelatus monticola Guignot, 1951
Copelatus montivagus Young, 1942
Copelatus mulangensis Bameul, 2003
Copelatus mundus Sharp, 1882
Copelatus mutabilis Bilardo & Rocchi, 1999
Copelatus mvoungensis Bilardo & Rocchi, 2004
Copelatus mysorensis Vazirani, 1970
Copelatus nakamurai Guéorguiev, 1970
Copelatus nangaensis Guignot, 1952
Copelatus nauclerus Guignot, 1939
Copelatus neavei J.Balfour-Browne, 1950
Copelatus neelumae Vazirani, 1973
Copelatus neoguineensis Zimmermann, 1919
Copelatus nigricans Sharp, 1882
Copelatus nigrolineatus Sharp, 1882
Copelatus nigropennis Zimmermann, 1927
Copelatus nigrostriatus Régimbart, 1895
Copelatus nilssoni Wewalka, 1982
Copelatus nimbaensis Legros, 1958
Copelatus nitens Bilardo & Rocchi, 1999
Copelatus nitidus Sharp, 1882
Copelatus nodieri Régimbart, 1895
Copelatus normalis Erichson, 1847
Copelatus notius Omer-Cooper, 1965
Copelatus nzei Bilardo & Rocchi, 1999
Copelatus oblitus Sharp, 1882
Copelatus obscurus Sharp, 1882
Copelatus occultus Bilardo & Rocchi, 1995
Copelatus ogasawarensis Kamiya, 1932
Copelatus onorei Pederzani & Rocchi, 1982
Copelatus ornatipennis Zimmermann, 1926
Copelatus owas Régimbart, 1895
Copelatus pachys Guignot, 1955
Copelatus pallidus Régimbart, 1895
Copelatus paludorum Guignot, 1952
Copelatus pandanorum Scott, 1912
Copelatus pantosi Guignot, 1958
Copelatus papuensis J.Balfour-Browne, 1939
Copelatus parabaptus Guignot, 1955
Copelatus parallelipipedus Régimbart, 1895
Copelatus parallelus Zimmermann, 1920
Copelatus pardii Rocchi, 1990
Copelatus parisii Guignot, 1959
Copelatus parumstriatus Gschwendtner, 1934
Copelatus paryphes Guignot, 1955
Copelatus pederzanii Bilardo & Rocchi, 1995
Copelatus pereirai Guignot, 1955
Copelatus peridinus Guignot, 1955
Copelatus pinnifer Guignot, 1952
Copelatus piriensis Omer-Cooper, 1965
Copelatus platynotus Régimbart, 1895
Copelatus polystrigus Sharp, 1882
Copelatus ponomarenkoi Riha, 1974
Copelatus portior Guignot, 1956
Copelatus posticatus (Fabricius, 1801)
Copelatus poungai Bilardo & Rocchi, 2006
Copelatus predaveterus K.B.Miller, 2003
Copelatus prolixus Sharp, 1882
Copelatus prolongatus Sharp, 1882
Copelatus propino Guignot, 1960
Copelatus propinquus Régimbart, 1895
Copelatus proximus Sharp, 1882
Copelatus pulchellus (Klug, 1834)
Copelatus pulicarius Régimbart, 1895
Copelatus pumilus Régimbart, 1895
Copelatus punctatus Bilardo & Rocchi, 1995
Copelatus punctulatus Aubé, 1838
Copelatus quadrisignatus Régimbart, 1877
Copelatus racenisi Guignot, 1951
Copelatus ragazzii Régimbart, 1887
Copelatus regimbarti Branden, 1885
Copelatus restrictus Sharp, 1882
Copelatus rimosus Guignot, 1952
Copelatus rivalis Guignot, 1952
Copelatus rocchii Bilardo, 1982
Copelatus romani Zimmermann, 1924
Copelatus royi Legros, 1958
Copelatus rubiginosus Guignot, 1954
Copelatus ruficapillus Régimbart, 1895
Copelatus ruteri Legros, 1954
Copelatus saegeri Guignot, 1955
Copelatus sahlbergi J.Balfour-Browne, 1939
Copelatus sallaei Sharp, 1882
Copelatus sanfilippoi Bilardo & Pederzani, 1972
Copelatus scalptus Guignot, 1954
Copelatus scaphites Guignot, 1955
Copelatus schaefferi Young, 1942
Copelatus schereri Wewalka, 1981
Copelatus schuhi Hendrich & Balke, 1998
Copelatus scutatus Guignot, 1952
Copelatus scytalotus Guignot, 1956
Copelatus senegalensis Lagar & Hernando, 1991
Copelatus sexstriatus Sharp, 1882
Copelatus sharpi Branden, 1885
Copelatus silvestrii Régimbart, 1903
Copelatus simoni Régimbart, 1889
Copelatus singularis Bilardo & Rocchi, 1995
Copelatus sociennus J.Balfour-Browne, 1952
Copelatus solitarius Sharp, 1882
Copelatus sordidipennis Régimbart, 1895
Copelatus spangleri Vazirani, 1974
Copelatus speciosus Régimbart, 1892
Copelatus spoliatus Guignot, 1955
Copelatus stavropolitanus Riha, 1974
Copelatus stillicidicola Bilardo & Rocchi, 1995
Copelatus striatellus Boheman, 1848
Copelatus striaticollis Lucas, 1857
Copelatus striatopterus Aubé, 1838
Copelatus striatulus Aubé, 1838
Copelatus strigipennis (Laporte, 1835)
Copelatus strigosulus (Fairmaire, 1878)
Copelatus strinatii Guignot, 1958
Copelatus striolatus Peschet, 1917
Copelatus stygis Guignot, 1958
Copelatus subdeficiens Régimbart, 1902
Copelatus subsimilis Guignot, 1958
Copelatus substriatus Kirsch, 1873
Copelatus subterraneus Guéorguiev, 1978
Copelatus sudrei Bameul, 2003
Copelatus sulcatus Sharp, 1882
Copelatus sulcipennis (Laporte, 1835)
Copelatus sumbawensis Régimbart, 1899
Copelatus suppar Guignot, 1956
Copelatus supplementaris Régimbart, 1895
Copelatus sylvaticus Guignot, 1952
Copelatus takakurai Satô, 1985
Copelatus tanaus Guignot, 1953
Copelatus taprobanicus Wewalka & Vazirani, 1985
Copelatus tenebrosus Régimbart, 1880
Copelatus teranishii Kamiya, 1938
Copelatus terminalis Sharp, 1882
Copelatus ternatensis Régimbart, 1899
Copelatus thiriouxi Peschet, 1917
Copelatus thrasys Guignot, 1952
Copelatus tibialis Sharp, 1882
Copelatus tinctor Guignot, 1954
Copelatus togoensis Régimbart, 1895
Copelatus tomokunii Satô, 1985
Copelatus tostus J.Balfour-Browne, 1950
Copelatus trifilis Guignot, 1958
Copelatus triglyphus Guignot, 1955
Copelatus trilobatus Régimbart, 1895
Copelatus tschaga Bilardo & Pederzani, 1972
Copelatus tucuchiensis J.Balfour-Browne, 1939
Copelatus tulagicus Guignot, 1942
Copelatus uludanuensis Hendrich & Balke, 1995
Copelatus undecimstriatus Aubé, 1838
Copelatus unguicularis Régimbart, 1903
Copelatus usagarensis Zimmermann, 1926
Copelatus vagatus Guignot, 1952
Copelatus vagestriatus Zimmermann, 1919
Copelatus vagus Bilardo & Rocchi, 1995
Copelatus validus Sharp, 1882
Copelatus vanninii Bilardo & Rocchi, 1999
Copelatus variegatus Régimbart, 1895
Copelatus venustus Bilardo & Rocchi, 1995
Copelatus vigintistriatus Fairmaire, 1869
Copelatus vigintisulcatus Régimbart, 1895
Copelatus villiersi Guignot, 1950
Copelatus virungaensis Bilardo, 1982
Copelatus vitraci Legros, 1948
Copelatus vivax Guignot, 1953
Copelatus wallacei J.Balfour-Browne, 1939
Copelatus waltoni J.Balfour-Browne, 1950
Copelatus wewalkai Holmen & Vazirani, 1990
Copelatus weymarni J.Balfour-Browne, 1947
Copelatus xanthocephalus Régimbart, 1899
Copelatus yacumensis Guignot, 1957
Copelatus zadiensis Bilardo & Rocchi, 1995
Copelatus zimmermanni Gschwendtner, 1934

References

Miller,K.B, Blake,M. The unusual occurrence of aquatic beetles in amber, copelatus aphroditae balke, n. sp. and C. predaveterus miller, n. sp., (Coleoptera: Dytiscidae: Copelatinae). Proc. of the Entomological Society of Washington, 2003, vol. 105, no4, pp. 809–815
Balke,M et al., MtDNA phylogeny and biogeography of Copelatinae, a highly diverse group of tropical diving beetles (Dytiscidae). Molecular Phylogenetics and Evolution Vol. 32, Issue 3, 2004
Dytiscidae Species List at Joel Hallan's Biology Catalog. Texas A&M University. Retrieved on 7 May 2012.
Caetano, David S., de C. Bena, Daniela, & Vanin, Sergio A. "Copelatus cessaima sp. nov. (Coleoptera: Dysticidae: Copelatinae): first record of a troglomorphic diving beetle from Brazil." Zootaxa 3710 (3): 226–232.
Megna, Y.S.; Epler, J.H. 2012: A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae). Acta Entomologica Musei Nationalis Pragae, 52(2): 383–410.
Megna, Y.S.; Epler, J.H. 2012: A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae). Acta Entomologica Musei Nationalis Pragae, 52(2): 383–410.