Ascalaphini

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Neuropterida
Ordo: Neuroptera
Subordo: Myrmeleontiformia
Superfamilia: Myrmeleontoidea

Familia: Myrmeleontidae
Subfamilia: Ascalaphinae
Tribus: Ascalaphini
Genera: Abascalaphus – Acheron – Acmonotus – Afroasca – Agadirius – Agrionosoma – Angolania – Angustacsa – Ascalaphodes – Ascalaphus – Ascalohybris – Ascalohybris – Ascapseudoptynx – Aspoeckiella – Botjederinus – Brevibarbis – Bubomyiella – Bubopsis – Cirrops – Deleproctophylla – Dentalacsa – Dicolpus – Disparomitus – Dixonotus – Dorsomitus – Encyoposis – Encyopsidius – Eremoides – Farakosius – Fillus – Forcepacsa – Glyptobasis – Helcopteryx – Horischema – Kimulodes – Libelloides – Lobalacsa – Mabiza – Maezous – Mansellacsa – Megacmonotus – Nephelasca – Nephoneura – Nousera – Ogcogaster – Parascalaphus – Parasuphalomitus – Perissoschema – Phalascusa – Pictacsa – Pilacmonotus – Proctarrelabis – Protacheron – Protidricerus – Protobubopsis – Pseudencyoposis – Pseudodisparomitus – Pseudohybris – Pseudoproctarrelabris – Puer – Siphlocerus – Stephanolasca – Strixomyia – Stylascalaphus – Suhpalacsa – Suphalomitus – Tytomyia – Ululomyia – Umbracsa – Venacsa
Name

Ascalaphini
References

Machado, R.J.P., Gilling, J.P., Winterton, S.L., Garzón-Orduña, I.J., Lemmon, A.R., Lemmon, E.M. & Oswald, J.D. 2018 (print 2019). Owlflies are derived antlions: anchored phylogenomics supports a new phylogeny and classification of Myrmeleontidae (Neuroptera). Systematic Entomology 44(2): 1–33. DOI: 10.1111/syen.12334 Paywall. ResearchGate Open access. Reference page.

Ascalaphini is the type subfamily of the neuropteran owlfly family. Most species are found in the tropics. Their characteristic apomorphy , shared with the Ululodini, is the ridge which divides each of their large compound eyes; both groups are thus sometimes known as split-eyed owlflies. The group has been alternatly treated as a subfamily of Ascalaphidae, when the family is treated separate from Myrmelontidae,[1] or as a tribe, when the ascalaphids are treated as a subfamily in an expanded Myrmelontidae.[2]

Like the other owlflies, they are insectivores. Imagines are cumbersome fliers and lack the strong mouthparts of dragonflies (which owlflies resemble at first glance, despite being not at all closely related insects) or other decidedly predatory insects, they are restricted to small and defenseless prey. The larvae on the other hand resemble antlions in appearance and habits and are voracious ambush predators, able to tackle prey like ants that will not be eaten without a struggle.

The ascalaphine split-eyed owlflies form one of the two main lineages of living Ascalaphidae, the other being the Haplogleniini which have unsplit eyes like their ancestors. The first fossil record of the Ascalaphinae dates to the Miocene, and the subfamily thus probably evolved in the latter half or towards the end of the Paleogene.[3]
Genera

There are 70 described genera in the subfamily Ascalaphinae.[2]
Ascalaphus sinister male
Glyptobasis sp. male

Abascalaphus Tjeder & Hansson, 1992
Acheron (insect) Lefèbvre 1842
Acmonotus McLachlan, 1871
Agadirius Badano & Pantaleoni, 2012
Agrionosoma van der Weele, 1909
Angolania Koçak & Kemal, 2008
Angustacsa New, 1984
Ascalaphodes McLachlan, 1871
Ascalaphus Fabricius, 1775
Ascalohybris Sziráki, 1998
Ascapseudoptynx Abrahám & Mészáros, 2006
Aspoeckiella Hölzel 2004
Botjederinus Abrahám, 2011
Brevibarbis Tjeder & Hansson, 1992
Bubomyiella Tjeder & Hansson, 1992
Bubopsis McLachlan 1898
Cirrops Tjeder 1980
Deleproctophylla Lefèbvrem 1842
Dentalacsa New, 1984
Dicolpus Gerstaecker, 1885
Disparomitus van der Weele, 1909
Dixonotus Kimmins, 1950
Dorsomitus Michel & Tjeder, 2018
Encyoposis McLachlan 1871
Encyopsidius Navás 1912
Eremoides Tjeder 1992
Farakosius Michel 1998
Fillus Navás 1919
Forcepacsa New 1984
Glyptobasis McLachlan 1871
Helcopteryx McLachlan 1871
Horischema Mészáros & Abrahám, 2003
Kimulodes Tjeder & Hansson 1992
Libelloides Schäffer 1763
Lobalacsa New 1984
Mabiza Tjeder & Hansson 1992
Maezous Ábrahám 2008
Mansellacsa Hölzel 2004
Megacmonotus New 1984
Nagacta Navás 1914
Nanomitus Navás 1912
Nephelasca Navás 1914
Nephoneura McLachlan 1871
Nousera Navás 1923
Ogcogaster Westwood 1847
Parascalaphus Martynova 1926
Parasuphalomitus New 1984
Perissoschema Mészáros & Abrahám 2003
Phalascusa Kolbe 1897
Pictacsa New 1984
Pilacmonotus New 1984
Proctarrelabis Lefèbvre 1842
Protacheron Weele 1909
Protidricerus Weele 1909
Protobubopsis van der Weele 1909
Pseudencyoposis van der Weele, 1909
Pseudodisparomitus New 1984
Pseudohybris van der Weele 1909
Pseudoproctarrelabris van der Weele 1909
Puer (insect) Lefèbvre 1842
Siphlocerus McLachlan 1871
Stephanolasca van der Weele 1909
Strixomyia Tjeder 1989
Stylascalaphus Sziráki 1998
Suhpalacsa Lefèbvre, 1842
Suphalomitus van der Weele 1909
Tytomyia Tjeder & Hansson 1992
Ululomyia Tjeder 1992
Umbracsa New 1984
Venacsa New 1984

References

Jones, Joshua R. (6 October 2019). "Total‐evidence phylogeny of the owlflies (Neuroptera, Ascalaphidae) supports a new higher‐level classification". Zoologica Scripta. 48 (6): 761–782. doi:10.1111/zsc.12382.
Machado, R. J. P.; Gillung, J. P.; Winterton, S. L.; Garzon‐Orduña, I. J.; Lemmon, A. R.; Lemmon, E. M.; Oswald, J. D. (2018). "Owlflies are derived antlions: Anchored phylogenomics supports a new phylogeny and classification of Myrmeleontidae (Neuroptera)". Systematic Entomology. 44 (2): 418–450. doi:10.1111/syen.12334.
See references in Haaramo (2008)