Aphodiinae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Aphodiinae
Tribus (11): Aphodiini - Corythoderini - Eupariini - Odochilini - Odontolochini - Phycochini - Psammodiini - Rhyparini - Stereomerini - Termitoderini - Thinorycterini

[source: Smith (2006)]

Genera (361): Acanthaphodius – Acanthobodilus – Acrossidius – Acrossoides – Acrossus – Adebrattia – Adeloparius – Afroataenius – Afrodiapterna – Afrodiastictus – Aganocrossus – Agoliinus – Agolius – Agrilinellus – Agrilinus – Ahermodontus – Aidophus – Airapus – Alloblackburneus – Allobodilus – Alocoderus – Amerilochus – Amidorus – Ammoecioides – Ammoecius – Anaetius – Annegialia – Anomalurobius – Anomius – Aparammoecius – Aphodaulacus – Aphodiellus – Aphodiopsis – Aphodius – Aphodobius – Aphodoharmogaster – Aphodopsammobius – Aphotaenius – Apsteiniella – Arupaia – Aschnarhyparus – Ataeniopsis – Ataenius – Australagolius – Australammoecius – Australaphodius – Australoxenella – Ballucus – Basilewskyanus – Batesiana – Biralus – Blackburneus – Bobiricola – Bodiloides – Bodilopsis – Bodilus – Bordatius – Boucomontiellus – Brachiaphodius – Brindalus – Bruchaphodius – Bruneixenus – Calamosternus – Caligodorus – Calocolobopterus – Candezeollus – Carinaulus – Cartwrightia – Cephalocyclus – Cesamexico – Chaetopisthes – Cheleion – Chilothorax – Chittius – Cinacanthus – Cnemargulus – Cnematoplatys – Cnemisus – Coelotrachelus – Colobopteridius – Colobopteroides – Colobopterus – Coprimorphus – Coptochiroides – Coptochirus – Corythoderus – Craterocyphus – Cryptaphodius – Cryptoscatomaseter – Daintreeola – Danielssonia – Dellacasiellus – Dialytellus – Dialytes – Dialytodius – Diapterna – Diastictus – Didactylia – Dilortomaeus – Doaphius – Drepanocanthoides – Drepanocanthus – Dudleyellus – Emadiellus – Eocorythoderus – Erytodes – Erytus – Esymus – Eudolus – Euhemicyclium – Euheptaulacus – Euorodalus – Euotophorus – Euparia – Euparixia – Euparixoides – Euparotrix – Eupleurus – Exaphodius – Exoxyomus – Ferrerianus – Flaviellus – Flechtmanniella – Geomyphilus – Geopsammodius – Gilletianus – Goiginus – Gonaphodiellus – Gonaphodioides – Gonaphodiopsis – Gonaphodius – Gongrolophus – Gordonius – Grandinaphodius – Granulopsammodius – Guanyinaphodius – Hadrorhyparus – Harmodactylus – Harmogaster – Haroldaphodius – Haroldiataenius – Haroldiellus – Hemicyclium – Hemycorythoderus – Heptaulacus – Hornietus – Hornosus – Iarupea – Iguazua – Ingogius – Irrasinus – Jalisco – Koreoxyomus – Koshantschikovius – Labarrus – Lechorodius – Leiopsammodius – Leptorhyparus – Limarus – Linargius – Lindbergianus – Lioglyptoxenus – Liothorax – Loboparius – Lomanoxia – Lomanoxoides – Longaphodius – Loraphodius – Loraspis – Lorditomaeus – Lunaphodius – Luxolinus – Macroretroides – Macroretrus – Maculaphodius – Martineziana – Mecynodes – Megateloides – Megatelus – Melinopterus – Mendidaphodius – Mendidius – Merogyrus – Mesontoplatys – Messyrhus – Microtermitodius – Microteuchestes – Monteitheolus – Mothon – Mozartius – Myrhessus – Myrmecaphodius – Mysarus – Nanotermitodius – Napoa – Neagolius – Neocalaphodius – Neocolobopterus – Neodiapterna – Neoemadiellus – Neoheptaulacus – Neopsammodius – Neorhyssemus – Neotrichaphodioides – Neotrichiorhyssemus – Neotrichonotulus – Nettelislasia – Nialaphodius – Nialosternus – Nialus – Nimbus – Nipponaphodius – Nipponoagoliinus – Nobiellus – Nobius – Nolicus – Notocaulus – Obaphodius – Odochilus – Odontacrossus – Odontolochus – Odontolytes – Odontopsammodius – Odontoxyomus – Orammoecius – Orocanthus – Orodaliscoides – Orodaliscus – Oromus – Oscarinus – Osmanius – Otophorus – Oxyataenius – Oxycorythus – Oxyomoides – Oxyomus – Ozodius – Parabodilus – Paracoptochirus – Paracorythoderus – Paracrossidius – Paradeloparius – Paradidactylia – Parammoecius – Paranimbus – Paraplesiataenius – Parapsammodius – Pararhyssemus – Parataenius – Paratrichaphodius – Pardalosus – Passaliolla – Paulianellus – Petrovitzius – Phaeaphodius – Phalacronothus – Pharaphodius – Pholeoaphodius – Phycochus – Pittinius – Plagiogonus – Planolinellus – Planolinoides – Planolinus – Platyderides – Platytomus – Pleuraphodius – Pleurophorus – Podotenoides – Podotenus – Proctophanes – Psammodaphodius – Psammodius – Psammorpha – Pseudacrossus – Pseudagolius – Pseudataenius – Pseuderytus – Pseudesymus – Pseudoagoliinus – Pseudoahermodontus – Pseudocoelotrachelus – Pseudodrepanocanthus – Pseudogonaphodiellus – Pseudoheptaulacus – Pseudomothon – Pseudopharaphodius – Pseudopodotenus – Pseudosaprosites – Pseudostereomera – Pseudotetraclipeoides – Pseudoteuchestes – Pseudoxyomus – Pterobius – Pubinus – Qingaphodius – Rakovicius – Renaudius – Rhinocerotopsis – Rhodaphodius – Rhyparus – Rhyssemodes – Rhyssemorphus – Rhyssemus – Rugaphodius – Sahlbergianus – Saprolochus – Saprositellus – Saprosites – Saprovisca – Sariangus – Scabrostomus – Schaefferellus – Selviria – Serraphodius – Setodius – Setylaides – Siamaphodius – Sicardia – Sigorus – Simogonius – Sinaphodius – Sinodiapterna – Sitiphus – Skelleyanus – Stebnickiella – Stenotothorax – Stereomera – Strigodius – Subrinus – Sugrames – Supertermitoderus – Sussorca – Sybacodes – Sybax – Symphodon – Tanyana – Termitaxis – Termitoderus – Termitodiellus – Termitodius – Termitopisthes – Tesarius – Tetraclipeoides – Teuchestes – Thinorycter – Trichaphodiellus – Trichaphodioides – Trichaphodius – Trichiopsammobius – Trichiorhyssemus – Trichonotuloides – Trichonotulus – Trigonoscelus – Tristaphodius – Turanella – Vladimirellus – Volinus – Xenoheptaulacus – Xeropsamobeus – Youngaphodius
Name

Aphodiinae Leach, 1815

Type genus: Aphodius Illiger, 1798 [stem = Aphodi-]

References
Additional references

Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204. DOI: 10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2 PDF
Stebnicka, Z.T. 2009: Aphodiinae of Australia (Coleoptera: Scarabaeidae). Iconography. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Kraków. ISBN 978-83-61358-24-4 [not seen]

Links

Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on April 20, 2017]

Vernacular names
English: Aphodiine dung beetle
日本語: マグソコガネ亜科
русский: Афодиины
svenska: Dyngbaggar

Aphodiinae is a subfamily of the scarab beetle family, Scarabaeidae. Members of this subfamily are known commonly as the small dung beetles and many, but not all, are dung beetles.[1] These beetles are found worldwide.[1]

These beetles are small scarab beetles, most less than 8 millimeters long. Many have small mandibles that are covered by a widened clypeus, the exoskeleton plate above the mouth. The feet are clawed.[1]

This is a diverse subfamily with varied life strategies and habitat types. Many species are dung beetles, which collect and feed on animal dung. Other species are detritivores or saprophages, which feed on dead matter, and some are predatory. Some are known as inquilines, living in ant or termite nests, and some are sand-dwelling beetles.[1] A survey of South American aphodiines found them in diverse habitat types including temperate rainforests, high-elevation Andean grassland, Patagonian steppe, coastal sand dunes, and subantarctic beech forest.[2]

There is not full agreement on the taxonomy of the subfamily, but some classification schemes divide it into 11 tribes with about 280 genera and a total of about 3200 species worldwide.[1]
Tribes

Aphodiini
Corythoderini
Didactyliini
Eupariini
Odochilini
Odontolochini
Proctophanini
Psammodiini
Rhyparini
Stereomerini
Termitoderini

References

Skelley, P. E. Aphodiinae. Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2008 Version.
Smith, A. B. T. and P. E. Skelley. (2007). A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America. Zootaxa 1458.