Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Crustacea
Superclassis: Multicrustacea
Classis: Malacostraca
Subclassis: Eumalacostraca
Superordo: Peracarida
Ordo: Isopoda
Cladus: Scutocoxifera
Subordo: Cymothoida
Superfamilia: Cymothooidea
Familia: Cymothoidae
Genus: Ceratothoa
Species: C. angulata – C. argus – C. atherinae – C. banksii – C. barracuda – C. capri – C. carinata – C. collaris – C. contracta – C. gaudichaudii – C. gilberti – C. gobii – C. guttata – C. hemirhamphi – C. huttoni – C. imbricata – C. italica – C. laticauda – C. marisrubri – C. novae-zelandiae – C. oestroides – C. oxyrrhynchaena – C. parallela – C. parva – C. potassoniensis – C. retusa – C. steindachneri – C. transversa – C. trigonocephala – C. trilles – C. usacarangis – C. verrucosa
Add (2): C. africanae – C. famosa
References
Hadfield, K.A., Bruce, N.L. & Smit, N.J. 2016. Redescription of poorly known species of Ceratothoa Dana, 1852 (Crustacea, Isopoda, Cymothoidae), based on original type material. ZooKeys 592: 1–53. DOI: 10.3897/zookeys.592.8098. Reference page.
Horton, T., 2000: Ceratothoa steindachneri (Isopoda: Cymothoidae) new to British waters with a key to north-east Atlantic and Mediterranean Ceratothoa. J. Mar. Biol. Ass. U.K., 80: 1041–1052. Full article: [1].
Martin, M.B.; Bruce, N.L.; Nowak, B.F. 2013: Redescription of Ceratothoa carinata (Bianconi, 1869) and Ceratothoa oxyrrhynchaena Koelbel, 1878 (Crustacea: Isopoda: Cymothoidae), buccal-attaching fish parasites new to Australia. Zootaxa 3683(4): 395–410. DOI: 10.11646/zootaxa.3683.4.4 Reference page.
Martin, M.B., Bruce, N.L. & Nowak, B.F. 2015: Review of the fish-parasitic genus Ceratothoa Dana, 1852 (Crustacea: Isopoda: Cymothoidae) from Australia, with description of two new species. Zootaxa 3963(3): 251–294. DOI: 10.11646/zootaxa.3963.3.1. Preview (PDF) Reference page.
van der Wal, S., Haug, J.T. 2020. Shape of attachment structures in parasitic isopodan crustaceans: the influence of attachment site and ontogeny. PeerJ. 8:e9181. DOI: 10.7717/peerj.9181 Open access Reference page.
Vernacular names
日本語: ヒゲブトウオノエ属
Ceratothoa is a genus of isopod ectoparasites of teleost fish,[2][3] first described by James Dwight Dana in 1852.[1] Infection by Ceratothoa can cause anaemia, lesions, growth retardation, emaciation, and mortality in their fish hosts.[3]
Species
Currently, 26 valid species in this genus are recognized:[1]
Ceratothoa africanae Hadfield, Bruce & Smit, 2014[4]
Ceratothoa angulata (Richardson, 1910)
Ceratothoa arimae (Nunomura, 2001)
Ceratothoa banksii (Leach, 1818)
Ceratothoa barracuda Martin, Bruce & Nowak, 2015
Ceratothoa capri (Trilles, 1964)
Ceratothoa carinata (Bianconi, 1869)
Ceratothoa collaris Schiödte & Meinert, 1883
Ceratothoa famosa Hadfield, Bruce & Smit, 2014[4]
Ceratothoa gilberti (Richardson, 1904)
Ceratothoa globulus Martin, Bruce & Nowak, 2015
Ceratothoa gobii Schiödte & Meinert, 1883
Ceratothoa guttata (Richardson, 1910)
Ceratothoa imbricata (Fabricius, 1775)
Ceratothoa italica Schiödte & Meinert, 1883
Ceratothoa marisrubri Trilles, Colorni & Golani, 1999
Ceratothoa oestroides (Risso, 1826)
Ceratothoa oxyrrhynchaena Koelbel, 1878
Ceratothoa parallela (Otto, 1828)
Ceratothoa retusa (Schioedte & Meinert, 1883)
Ceratothoa springbok Hadfield & Smit, 2020
Ceratothoa steindachneri Koelbel, 1879
Ceratothoa toyamaensis (Nunomura, 1993)
Ceratothoa trigonocephala (Leach, 1818)
Ceratothoa usacarangis (Avdeev, 1979)
Ceratothoa verrucosa (Schioedte & Meinert, 1883)
C. carinata and C. oxyrrhynchaena were redescribed in 2013,[5] and C. angulata, C. capri, C. carinata, C. collaris, C. gilberti, C. gobii, C. guttata, C. italica, C. oestroides, and C. verrucosa in 2016.[6]
A number of taxa are species inquirendae:[1]
Ceratothoa contracta (Miers, 1880)
Ceratothoa deplanata Bovallius, 1885
Ceratothoa gaudichaudii (H. Milne Edwards, 1840) (previously known as Ceratothoa rapax Heller, 1865)[7]
Ceratothoa novaezelandiae Filhol, 1885
Ceratothoa transversa (Richardson, 1901)
Ceratothoa triglae Gourret, 1891
References
Schotte M, Boyko CB, Bruce NL, Poore GC, Taiti S, Wilson GD, eds. (2021). "Ceratothoa Dana, 1852". World Marine, Freshwater and Terrestrial Isopod Crustaceans database. World Register of Marine Species. Retrieved 2 September 2021.
Horton, Tammy (2000). "Ceratothoa steindachneri (Isopoda: Cymothoidae) new to British waters with a key to north-east Atlantic and Mediterranean Ceratothoa". Journal of the Marine Biological Association of the United Kingdom. 80 (6): 1041–1052. doi:10.1017/S0025315400003106.
Horton, Tammy; Diamant, Arik; Galil, Bella (2004). "Ceratothoa steindachneri (Isopoda, Cymothoidae): An unusual record from the Mediterranean". Crustaceana. 77 (9): 1145–1148. doi:10.1163/1568540042900231.
Hadfield, Kerry; Bruce, Niel; Smit, Nico (2014). "Review of the fish parasitic genus Ceratothoa Dana, 1852 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of two new species". ZooKeys (400): 1–42. doi:10.3897/zookeys.400.6878. PMID 24843254. S2CID 6067384.
Martin, Melissa B.; Bruce, Niel L.; Nowak, Barbara F. (2013). "Redescription of Ceratothoa carinata (Bianconi, 1869) and Ceratothoa oxyrrhynchaena Koelbel, 1878 (Crustacea: Isopoda: Cymothoidae), buccal-attaching fish parasites new to Australia". Zootaxa. 3683 (4): 395–410. doi:10.11646/zootaxa.3683.4.4. PMID 25250460.
Hadfield, Kerry A.; Bruce, Niel L.; Smit, Nico J. (2016). "Redescription of poorly known species of Ceratothoa Dana, 1852 (Crustacea, Isopoda, Cymothoidae), based on original type material". ZooKeys. 592: 39–91. doi:10.3897/zookeys.592.8098. PMC 4926636. PMID 27408544.
Brusca, Richard C. (1981). "A monograph on the Isopoda Cymothoidae (Crustacea) of the eastern Pacific". Zoological Journal of the Linnean Society. 73 (2): 117–199. doi:10.1111/j.1096-3642.1981.tb01592.x.
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