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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Ordo: Cornales

Familia: Hydrostachyaceae
Genus: Hydrostachys
Species: H. angustisecta – H. bismarckii – H. decaryi – H. distichophylla – H. fimbriata – H. flabellifera – H. imbricata – H. insignis – H. longifida – H. lufirensis – H. lukungensis – H. maximus – H. monoicus – H. multifida – H. myriophylla – H. perrieri – H. plumosa – H. polymorpha – H. stolonifera – H. triaxialis – H. trifaria – H. verruculosa
Source(s) of checklist:
Name

Hydrostachys Thouars, 1806

Typus: Hydrostachys verruculosa A. Juss. LT designated by Bullock, Kew Bull. 14: 42 (1960)

Distribution
Native distribution areas:

Continental: Africa
Regional: Tropical Africa
Angola, KwaZulu-Natal, Madagascar, Malawi, Mozambique, Namibia, Tanzania, Zambia, Zaïre, Zimbabwe

References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition
References
Primary references

Thouars, L.M.A. du Petit, 1806. Gen. Nov. Madagasc.: 2
Additional references

Govaerts, R., Nic Lughadha, E., Black, N., Turner, R. & Paton, A. 2021. The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. Scientific Data 8(215): 1–10. DOI: 10.1038/s41597-021-00997-6 Reference page.

Links

Govaerts, R. et al. 2021. Hydrostachys in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2021 Oct 25. Reference page.
International Plant Names Index. 2021. Hydrostachys. Published online. Accessed: Oct 25 2021.
Tropicos.org 2021. Hydrostachys. Missouri Botanical Garden. Published online. Accessed: 25 Oct 2021.
Hassler, M. 2021. Hydrostachys. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2021. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2021 Oct 25. Reference page.
USDA, ARS, Germplasm Resources Information Network. Hydrostachys in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 08-Apr-12.

Vernacular names
한국어: 히드로스타키스속
русский: Гидростахис


Hydrostachys is a genus of about 22 species[2] of flowering plants native to Madagascar and southern and central Africa. It is the only genus in the family Hydrostachyaceae. All species of Hydrostachys are aquatic, growing on rocks in fast-moving water. They have tuberous roots, usually pinnately compound leaves, and highly reduced flowers on dense spikes.[3]

Phylogenetic placement of Hydrostachys is particularly problematic. Due to its specialized aquatic morphology, it has often been grouped with other aquatic plants, such as the family Podostemaceae. However, embryological, floral, and other morphological characters do not support this placement, and molecular data suggest that Hydrostachys is related to taxa in the order Cornales.[3][4] Its position in Cornales is uncertain; it may be basal to the rest of the order, or fall within the family Hydrangeaceae.[3][5][6] It shares few morphological similarities with other Cornales.[3]
Species

(This list may be incomplete.)

Hydrostachys decaryi
Hydrostachys distichophylla
Hydrostachys fimbriata
Hydrostachys imbricata
Hydrostachys laciniata
Hydrostachys longifida
Hydrostachys maxima
Hydrostachys monoica
Hydrostachys multifida
Hydrostachys perrieri
Hydrostachys plumosa
Hydrostachys stolonifera
Hydrostachys trifaria
Hydrostachys verruculosa

References

Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III" (PDF). Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x. Retrieved 2013-07-06.
Christenhusz, M. J. M.; Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. 261 (3): 201–217. doi:10.11646/phytotaxa.261.3.1.
Albach, D. C., Soltis, D. E., Chase, M. W., and Soltis, P. S. (2001). Phylogenetic placement of the enigmatic angiosperm Hydrostachys. Taxon 50, 781-805.
Angiosperm Phylogeny Group (2016). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV". Botanical Journal of the Linnean Society. 181 (1): 1–20. doi:10.1111/boj.12385. ISSN 0024-4074.
Fan, C. Z., and Xiang, Q. Y. (2003). Phylogenetic analyses of Cornales based on 26S rRNA and combined 26S rDNA-matK-rbcL sequence data. American Journal of Botany 90, 1357-1372.
Kubitzki, K. (2004). Cornaceae. In The Families and Genera of Vascular Plants Volume 6: Flowering Plants: Dicotyledons: Celastrales, Oxidales, Rosales, Cornales, Ericales (Kubitzki, ed.). Springer-Verlag, New York.

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