Quercus alnifolia

Life-forms

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Rosids
Cladus: Eurosids I
Ordo: Fagales

Familia: Fagaceae
Genus: Quercus
Species: Quercus alnifolia
Name

Quercus alnifolia Poech, 1842

Synonyms

Heterotypic
Quercus cypria Jaub. & Spach, Ill. Pl. Orient. 1: 110 (1843).
Quercus alnifolia var. argentea Hadjik. & Hand, Willdenowia 36: 796 (2006).

Distribution
Native distribution areas:

Continental: Asia-Temperate
Regional: Western Asia
Cyprus

References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition
References
Primary references

Poech, J. 1842. Enumeratio Plantarum hucusque Cognitarum Insulae Cypri... Vindobonae 12.

Links

Govaerts, R. et al. 2021. Quercus alnifolia in World Checklist of Selected Plant Families. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2021 Jun 27. Reference page.
Govaerts, R. et al. 2021. Quercus alnifolia in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2021 Jun 27. Reference page.
International Plant Names Index. 2021. Quercus alnifolia. Published online. Accessed: Jun 27 2021.
Tropicos.org 2021. Quercus alnifolia. Missouri Botanical Garden. Published on the internet. Accessed: 2021 Jun 27.
Hassler, M. 2021. Quercus alnifolia. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2021. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2021 Jun 27. Reference page.
Hassler, M. 2021. World Plants. Synonymic Checklist and Distribution of the World Flora. . Quercus alnifolia. Accessed: 27 Jun 2021.
USDA, ARS, Germplasm Resources Information Network. Quercus alnifolia in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 07-Oct-06.

Vernacular names
magyar: Égerlevelű tölgy

Quercus alnifolia, commonly known as the golden oak, is an evergreen oak species of Cyprus. Its common English name refers to the golden coloured lower surface of its leaves. Quercus alnifolia belongs to the endemic flora of the island and it is confined to the igneous geological complex of the Troodos Mountains. In February 2006, the parliament of Cyprus selected the golden oak to be the country's national tree.[3]

Taxonomy
A golden oak shrub in a pine stand near Karvounas, Troodos Mountains

Quercus alnifolia belongs to section Cerris, as with much of the mediterranean evergreen oak species. Occasional hybridization with kermes oak (Quercus coccifera ssp. calliprinos) has been described.[4]
Description

The golden oak is a much branched evergreen shrub or small tree up to 10 m high. Due to its short stature (in relation to other oaks) it is sometimes referred to as the dwarf oak.[5]

Its leaves are simple, obovate to suborbicular, 1.5–6(–10) cm long, 1–5 (–8) cm wide, glabrous and shining dark green above and densely golden or brownish tomentose below, with serrate margins and raised nervation. Leave petioles are strong, 6–10 (–12) mm long and pilose. Flowers are unisexual; male catkins are greenish-yellow forming spreading or pendulous clusters at the tips of the branches; female are axillary, solitary or in groups of 2–3. Acorns are narrowly obovate or subcylindrical, usually tapering towards base, 2–2.5 cm long and 0.8–1.2 cm wide, with a woody endocarp and cupule with strongly recurved scales.[6]
Distribution and habitat

Quercus alnifolia is restricted to Troodos Massif, where it grows on igneous geological substrate at an altitude of 400 to 1800m.[7] It occupies dry habitats in association with Pinus brutia or forms dense maquis in mesic habitats, characterized by deep forest soils.[8]
Ecological importance and conservation status

Golden oak offers soil stabilisation against erosion due to its ability to colonize stony and rocky slopes. In its distribution area, Quercus alnifolia is the most important broadleaved species forming pure or mixed stands, within the conifer (Pinus brutia, Pinus nigra) dominated forests of Cyprus. Thick stands of Quercus alnifolia in mesic habitats, considerably modify the humidity conditions of the site and form forest soils with "mull" humus favouring the existence of sciophilous herbal species.

Like other members of the oak genus, Quercus alnifolia forms mutually beneficial, ectomycorrhizal associations with various fungi. A preliminary study in 2011, reported over 80 mycorrhizal fungi to be associated with the golden oak; the total number, however, is estimated to be much higher.[9]

Golden oak is protected by the forest law of Cyprus, while the habitat type "Scrub and low forest vegetation of Quercus alnifolia (9390)" is a priority habitat of Europe (directive 92/43/EEC).[10] Large forest expanses of the species have been proposed for inclusion in the Natura 2000 ecological network of the European Union.
References

Gorener, V. & Beech, E (2017). "Quercus alnifolia". The IUCN Red List of Threatened Species. IUCN. 208. e.T194053A2295358. doi:10.2305/IUCN.UK.2017-3.RLTS.T194053A2295358.en.
"Quercus alnifolia Poech". World Checklist of Selected Plant Families (WCSP). Royal Botanic Gardens, Kew – via The Plant List.
Leonidou, Leo (February 3, 2006). "A flower for Cyprus". Cyprus Mail. Retrieved 2007-12-30.[dead link]
Neophytou, Ch.; Palli G.; Aravanopoulos F.A. (2007). "Morphological differentiation and hybridization between Quercus alnifolia Poech and Quercus coccifera L. (Fagaceae) in Cyprus" (PDF). Silvae Genetica. 56 (6): 271–277. doi:10.1515/sg-2007-0038. S2CID 91154432. Archived from the original (PDF) on 2011-07-18. Retrieved 2008-03-01.
Ashmore, Wendy; Arthur Bernard Knapp (1999). Archaeologies of Landscape: Contemporary Perspectives. Blackwell. p. 237. ISBN 0-631-21106-3.
Meikle, R.D. (1985). Flora of Cyprus. London: Bentham Moxon Trust. pp. 1481–1482. ISBN 0-9504876-3-5.
Merlo, Maurizio (2005). Valuing Mediterranean Forests: Towards Total Economic Value. CABI Publishing. p. 217. ISBN 0-85199-480-6.
Barbéro, M.; Quézel P. (1979). "Contribution à l' étude des groupements forestiers de Chypre". Documents Phytosociologiques (IV): 9–34.
Loizides, M. (2011). Quercus alnifolia: The indigenous golden oak of Cyprus and its fungi. Field Mycology 12 (3): 81–88. doi:10.1016/j.fldmyc.2011.06.004.
"Council Directive 92/43/EEC of 21 May 1992 on the conservation of natural habitats and of wild fauna and flora". EUR-Lex. European Union law. 1992-07-22. Retrieved 2008-01-01.