Phedimus

Life-forms

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Ordo: Saxifragales

Familia: Crassulaceae
Subfamilia: Sempervivoideae
Tribus: Umbiliceae
Genus: Phedimus
Subgenera: P. subg. Aizoon – P. subg. Phedimus

Species: P. aizoon – P. floriferus – P. hsinganicus – P. hybridus – P. kamtschaticus – P. litoralis – P. latiovalifolium – P. middendorffianus – P. obtusifolius – P. odontophyllus – P. selskianus – P. sichotensis – P. sikokianus – P. spurius – P. stellatus – P. stevenianus – P. stoloniferus – P. takesimensis – P. zokuriensis
Name

Phedimus Raf. (1817)

Type species: P. stellatus (L.) Raf.

Synonyms

Homotypic
Asterosedum Grulich, Preslia 56: 38. 1984, nom. superfl.
Heterotypic
Aizopsis Grulich, Preslia 56: 35. 1984.
Type species: A. aizoon (L.) Grulich
Spathulata (Borissova) Á.Löve & D.Löve, Taxon 34: 163. 1985, nom. inval.
Type species: S. spurium Sibth. ex Nyman

References

Rafinesque-Schmaltz, C.S. 1817. Amer. Monthly Mag. & Crit. Rev. 1: 438.
Hassler, M. 2019. Phedimus. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2019. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2019 Aug. 30. Reference page.
Gontcharova, S.B. & Gontcharov, A.A. 2009. Molecular phylogeny and systematics of flowering plants of the family Crassulaceae DC. Molecular Biology 43(5): 794. DOI: 10.1134/S0026893309050112 PDF from ResearchGate Reference page.
Govaerts, R. et al. 2019. Phedimus in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2019 Aug. 30. Reference page.
International Plant Names Index. 2009. Phedimus. Published online. Accessed: 18 Jul 2009.
Thiede, J. & Eggli, U. 2007. Crassulaceae in Kubitzki, K., Bayer, C. & Stevens, P.F. (Eds.) Flowering plants : Eudicots ; Berberidopsidales, Buxales, Crossosomatales, Fabales p.p., Geraniales, Gunnerales, Myrtales p.p., Proteales, Saxifragales, Vitales, Zygophyllales, Clusiaceae Alliance, Passifloraceae Alliance, Dilleniaceae, Huaceae, Picramniaceae, Sabiaceae pp. 83–119 DOI: 10.1007/978-3-540-32219-1_12 ISBN 3540322140 Crassulaceae PDF in Research Gate Reference page.

Vernacular names
suomi: Aasianmaksaruohot
svenska: Fetbladssläktet

Phedimus is a genus of the succulent family Crassulaceae (stonecrop family), with about 18 species, distributed in eastern Europe and Asia. The genus is described with two subgenera, but one of these is also recognized as a separate genus, Aizopsis. Phedimus kamtschaticus (syn. Sedum kamtschaticum) is widely grown as an ornamental ground cover and has gained the Royal Horticultural Society's Award of Garden Merit.[2]

Description

Species are perennial or rarely annual herbs, usually glabrous stems, sometimes woody at base, from thin woody rhizome. Leaves decussate or alternate, with narrow base, and several hydathodes on lower face along margins. The flowering branches are erect or descending, with dense inflorescences exhibiting many flowered terminal pleiochasia (several buds come out at the same time). Flowers 4-7 parts, sepals usually unequal, petals free, usually spreading. The fruits are follicles, usually spreading and seeds costate-papillate (incompletely connate), but multipapillate in Phedimus selskianus.[3]

When treated as Phedimus sensu lato, i.e. including Aizopsis, subgenus Phedimus (Phedimus sensu strictu) has creeping and rooting stems, flowers white, pink, red or purplish and chromosome numbers x=5,6,7, while subgenus Aizoon has annual shoots often woody at base, emerging from woody rhizomes, flowers yellow, orange or reddish and x=8.[3]
Taxonomy

Phedimus is a genus in the family Crassulaceae, subfamily Sempervivoideae, tribe Umbiliceae, together with three other genera. It was segregated from the very large cosmopolitan and polyphyletic genus, Sedum, where it was variously considered a section or subgenus.[4] Within the tribe it is a sister group to Aizopsis. There, it is distinguished by flower colouration (pink or white flowers compared to the yellow flowers of Aizopsis), floral morphology, distribution (Eurasia vs. Asia) and chromosome numbers (x=14 vs. 16). Some authors include Aizopsis as a subgenus:[1][3]

Phedimus subgenus Phedimus c. 5 species
Phedimus subgenus Aizoon c. 13 species

Distribution

Subgenus Phedimus Mediterranean to Caucasus. Subgenus Aizoon China, Japan, Korea and central Siberia.[3]
References

Gontcharova et al 2006.
"Sedum kamtschaticum orange stonecrop". The Royal Horticultural Society. 2021. Retrieved 22 June 2021.
Thiede & Eggli 2007.

Ohba et al 2000.

Bibliography

Thiede, J; Eggli, U (2007). "Crassulaceae". In Kubitzki, Klaus (ed.). Berberidopsidales, Buxales, Crossosomatales, Fabales p.p., Geraniales, Gunnerales, Myrtales p.p., Proteales, Saxifragales, Vitales, Zygophyllales, Clusiaceae Alliance, Passifloraceae Alliance, Dilleniaceae, Huaceae, Picramniaceae, Sabiaceae. pp. 83–119. ISBN 978-3540322146. (full text at ResearchGate)
Gontcharova, S. B.; Artyukova, E. V.; Gontcharov, A. A. (June 2006). "Phylogenetic relationships among members of the subfamily Sedoideae (Crassulaceae) inferred from the ITS region sequences of nuclear rDNA" (PDF). Russian Journal of Genetics. 42 (6): 654–661. doi:10.1134/S102279540606010X. S2CID 33249840.
Grulich, Vit (1984). "Generic division of Sedoideae in Europe and the adjacent regions" (PDF). Preslia. 56: 29–45.
Ohba, Hideaki; Bartholomew, Bruce M.; Turland, Nicholas J.; Kunjun, Fu (2000). "New Combinations in Phedimus (Crassulaceae)". Novon. 10 (4): 400–402. doi:10.2307/3392995. JSTOR 3392995.