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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Ordo: Ranunculales

Familia: Berberidaceae
Subfamilia: Podophylloideae
Genus: Achlys
Species: A. californica – A. japonica – A. triphylla
Name

Achlys DC., 1821

Type species: Leontice triphylla Sm.

References

De Candolle, A.P. 1821. Regni Vegetabilis Systema Naturale 2: 35.

Links

Hassler, M. 2019. Achlys. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2019. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2019 Apr. 22. Reference page.
International Plant Names Index. 2019. Achlys. Published online. Accessed: Apr. 22 2019.
The Plant List 2013. Achlys in The Plant List Version 1.1. Published on the internet. Accessed: 2019 Apr. 22.
Tropicos.org 2019. Achlys. Missouri Botanical Garden. Published on the internet. Accessed: 2019 Apr. 22.

Vernacular names

Achlys is a small genus of flowering plants in the barberry family (Berberidaceae), which it shares with genera such as Berberis and Vancouveria.[1] It is named after the Greek goddess Achlys.[2]

Species

There are either two or three species, depending on the authority. Achlys triphylla and Achlys californica are both native to western North America. Another Achlys is found in Japan: some authorities treat this as a subspecies of A. triphylla, while others, especially in older treatments, call this Achlys japonica. Still others consider A. triphylla and A. californica too similar to be separate species. The common names for these plants include vanilla leaf (sometimes written as vanilla-leaf or vanillaleaf, depending on the taxonomist or flora), deer's foot and sweet after death, referring to the vanilla scent of its crushed leaves.[1][3]

The Plant List recognizes two species, with A. californica regarded as a subspecies of A. triphylla

Achlys triphylla (Sm.) DC. – British Columbia, Washington, Oregon, NW California[4]
Achlys japonica Maxim. – Japan

Covered with fine hairs, Achlys triphylla achenes are concave toward stem and fleshy.[3]
Description

Achlys triphylla, known in western North America as vanillaleaf, is an erect perennial plant that sprouts from a creeping rhizome. Leaves are long-petioled and palmately divided into three leaflets. Flowers are small and lack sepals and petals, but instead have long showy white stamens that form single erect spikes.[1] The leaflets give a great hint to the identification of the plant: bend back the middle leaflet and you have an upside-down set of moose antlers. Alternatively, bend back the two side leaflets and you have a goose or deer foot (hence the common name). In the Pacific Northwest, Achlys triphylla is ubiquitous in moist shady forests west of the Cascades at low to middle elevations from Vancouver island and southern British Columbia south to northern California.[3]

The plants are spaced widely on the rhizomes, but often overlap in large networks that result in carpets of Achlys that dominate the near-surface understory. Achlys seems to prefer moist soil, so at middle to higher elevations it is easier to find them along streambanks or well-shaded ravines.[3]
Insect repellent

When dried properly, the plants are strongly aromatic and smell of vanilla. Besides serving as an excellent tent air freshener, Achlys was used by native tribes of southern British Columbia as an insect repellent. The dried leaves were hung in bunches in doorways to ward off flies and mosquitoes, and it's not unheard of for naturalists to rub the dried or even fresh leaves on exposed skin when hiking the Olympics or Cascades during the summer mosquito season.[3]

Japanese Achlys are quite similar to those found in western North America.
References

Gilkey, Helen M. & Dennis, La Rea J. (2001). Handbook of Northwestern Plants. Oregon State University Press. ISBN 0-87071-490-2 p. 161
Gledhill, David (2008). The Names of Plants. Cambridge University. ISBN 9780521866453.
Pojar & MacKinnon (2004). Plants of the Pacific Northwest Coast (Revised Edition). Lone Pine Publishing. ISBN 1-55105-530-9 p. 312
Biota of North America Program 2013 county distribution map

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