Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
SuperOrdo: Eupulmonata
Ordo: Stylommatophora
SubOrdo: Helicina
InfraOrdo: Helicoidei
Superfamilia: Sagdoidea
Familia: Zachrysiidae
Genera (1): Zachrysia
Name
Zachrysiidae Robinson Sei & Rosenberg, 2017
References
Links
Zachrysiidae in the World Register of Marine Species
Zachrysiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod molluscs in the superfamily Sagdoidea.[1] It is native to Cuba, though some species have been introduced to Florida, West Indies islands, the Yucatán Peninsula of Mexico and Panama.[2]
Description
Zachrysiidae have solid and thick shells 16-70 mm in width and pale yellow to orange in colour. The shape is generally globose but depressed and somewhat eccentrically and unevenly coiled. The protoconch is 1.5 whorls that are polished and shiny with faint, microscopic, spiral lines. The teleoconch is 4-4.5 whorls and varies in sculpture from sharply and finely ribbed to almost smooth. There may be a microsculpture of closely crowded granules and these are sometimes arranged in irregular spiral striae. The aperture is oblique and downturned at maturity, with a thickened and reflexed lip. The baso-columellar margin is expanded with a slight tooth. Adults are not umbilicate and juveniles are minutely umbilicate.[2]
This family is unique in its reproductive anatomy. The right ommatophoral retractor muscle passes to the left of the genitalia. The phallus is thick and the phallus chamber in most species has one or two fleshy structures (stimulus or stimulators). The epiphallus is very short. The flagellum is thick and blunt-ended. Most species also have another structure, variously called the accessory flagellum or the penial caecum, attached separately to the phallus. The gametolytic gland duct is coiled around the phallus retractor muscle and separate from the oviduct throughout its length. Additionally, the gametolytic gland duct is short, swollen basally and has a relatively large reservoir. The ovotestis or gonad consists of very elongated acini, and there is no vagina.[2]
Classification
Zachrysia was once placed in the family Pleurodontidae in superfamily Helicoidea. A 2017 phylogenetic analysis transferred it to the new family Zachrysiidae in the superfamily Sagdoidea.[2]
Genera
Zachrysia Pilsbry, 1894
References
MolluscaBase eds. (2021). MolluscaBase. Zachrysiidae D. G. Robinson, Sei & Rosenberg, 2017. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=1057795 on 5 February 2021
Sei, Makiri; Robinson, David G; Geneva, Anthony J; Rosenberg, Gary (23 November 2017). "Doubled helix: Sagdoidea is the overlooked sister group of Helicoidea (Mollusca: Gastropoda: Pulmonata)". Biological Journal of the Linnean Society. 122 (4): 697–728. doi:10.1093/biolinnean/blx082. ISSN 0024-4066.
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