Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
SuperOrdo: Eupulmonata
Ordo: Stylommatophora
SubOrdo: Helicina
InfraOrdo: Orthalicoidei
Superfamilia: Orthalicoidea
Familia: Orthalicidae
Genera (14 + †1): Aposcutalus – Clathrorthalicus – Corona – Hemibulimus – Kara – Laeorthalicus – Liguus – Orthalicus – Paeniscutalus – Porphyrobaphe – Quechua – Scholvienia – Sultana – Trachyorthalicus – †Cortana
Name
Orthalicidae Martens, 1860
References
Breure A.S.H., 2009. New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World. Zootaxa 2065: 25–50. Abstract & excerpt
Breure, A.S.H.; Ablett, J.D. 2015. Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys, 470: 17–143. DOI: 10.3897/zookeys.470.8548 Open access. Reference page.
Breure A.S.H. & Borrero F.J., 2008. An annotated checklist of the land snail family Orthalicidae (Gastropoda: Pulmonata: Orthalicoidea) in Ecuador, with notes on the distribution of the mainland species. Zootaxa 1768: 1–40. Abstract & excerpt
Breure A.S.H. & Schlögl J., 2010. Additional notes on Orthalicidae from the Chimantá massif, Venezuelan Guayana, with descriptions of new species of Plekocheilus Guilding, 1828 (Mollusca: Gastropoda). Zootaxa 2416: 51–60. Preview
Links
Orthalicidae in the World Register of Marine Species
Orthalicidae (orthalicid land snails) are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the subfamily Orthalicoidea of the order Stylommatophora.[2]
They are medium-sized to large snails, from about 3 cm (about 1.2 inches) to 9 cm (about 3.5 inches) in shell length
Taxonomy
In former times, this family was often known as the Bulimulidae, but this term may also denote what today is the subfamily Bulimulinae. The subfamily Bulimulinae replaces the former family Bulimulidae Crosse & P. Fischer, 1873.
2005 taxonomy
The Orthalicidae belong to the Orthalicoidea, a superfamily in the order Sigmurethra. Like other stylommatophorans, the Sigmurethra belong to the suborder Helicina. Among the three subfamilies of Orthalicidae, the Bulimulinae are the most diverse, consisting of three tribes:[3]
subfamily Amphibuliminae P. Fischer, 1873 - synonym Peltellinae Gray, 1855 (check decision of International Commission on Zoological Nomenclature for Peltellinae) - 17 species[4]
subfamily Bulimulinae Tryon, 1867 - 1243 species[4]
tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
tribe Odontostomini Pilsbry & Vanatta, 1898 - 125 species[4]
tribe Simpulopsini Schileyko, 1999 - synonym: Tomogeridae Jousseaume, 1877
subfamily Orthalicinae Albers, 1850 - synonym: Liguidae Pilsbry, 1891 - 83 species[4]
2010 taxonomy
Breure et al. (2010)[5] elevated Bulimulinae to Bulimulidae, Odontostomini to Odontostomidae, Amphibuliminae to Amphibulimidae.[5]
2012 taxonomy
Breure & Romero (2012)[6] confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae.[6]
Genera
Genera within the family Orthalicidae include:
Aposcutalus Dutra & Leme, 1985
Clathrorthalicus Strebel, 1909
Corona Albers, 1850[7][6][8]
† Cortana Salvador & Simone, 2013
Hemibulimus Martens, 1885[8]
Kara Strebel, 1910[9]
Liguus Montfort, 1810
Orthalicus Beck, 1837 - type genus of the family Orthalicidae[3][6]
Paeniscutalus Wurtz, 1947
Porphyrobaphe Shuttleworth, 1856[6][8]
Quechua Strebel, 1910
Scholvienia Strebel, 1910
Sultana Shuttleworth, 1856[8]
References
Albers J. C. 1860. (Manuscript edited by Eduard von Martens) Die Heliceeen nach natürlicher Vervandtschaft systematisch geordnet. Edition 2, Engelmann, Leipzig, xviii + 359 pp., Orthalicidae is at page 209.
MolluscaBase eds. (2021). MolluscaBase. Orthalicidae Martens, 1860. Accessed through: World Register of Marine Species at: http://marinespecies.org/aphia.php?p=taxdetails&id=870044 on 2021-02-19
Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
Breure A., Groenenberg D. & Schilthuizen M. (2010). "Gondwana revisited: New insights in the phylogenetic relationships within the Orthalicoidea (Mollusca, Stylommatophora)". poster. PDF Archived 2012-03-09 at the Wayback Machine.
Breure A. S. H., Groenenberg D. S. J. & Schilthuizen M. (2010). "New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda, Stylommatophora) based on 28S sequence data". Basteria 74(1-3): 25-31.
Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. doi:10.1127/arch.moll/1869-0963/141/001-020.
Breure A. S. H. & Mogollón Avila V. (2010). "Well-known and little-known: miscellaneous notes on Peruvian Orthalicidae (Gastropoda, Stylommatophora)". Zoologische Mededelingen 84. HTM Archived 2018-09-26 at the Wayback Machine.
Mónica Lucía Vera Ardila (2008). "Lista de los géneros de moluscos terrestres de Colombia (Mollusca: Gastropoda: Prosobranchia: Mesogastropoda y Pulmonata: Stylommatophora)" (PDF). Biota Colombiana. 9 (1): 39–62.
Breure A. S. H. (2011) "Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species". ZooKeys 101: 1–50. doi:10.3897/zookeys.101.1133.
Further reading
(in German) Neubert E. & Janssen R. (2004). "Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae". Archiv für Molluskenkunde 133: 193–297. abstract.
Ramirez J. & Ramírez R. (2010). "Análisis de la estructura secundaria del LSU rRNA mitocondrial de caracoles terrestres peruanos (Orthalicidae: Gastropoda) / Analysis of the secondary structure of mitochondrial LSU rRNA of Peruvian land snails (Orthalicidae: Gastropoda)." Revista Peruana de Biología 17(1): 53–57. PDF.
Ramirez J., Ramírez R., Romero P., Chumbe A. & Ramírez P. (2009). "Posición evolutiva de caracoles terrestres peruanos (Orthalicidae) entre los Stylommatophora (Mollusca: Gastropoda) / Evolutionary position of Peruvian land snails (Orthalicidae) among Stylommatophora (Mollusca: Gastropoda)". Revista Peruana de Biología 16(1): 51–56. PDF.
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