Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
Superordo: Eupulmonata
Ordo: Ellobiida
Superfamilia: Ellobioidea
Familia: Ellobiidae
Subfamiliae (6): Carychiinae - Ellobiinae - Melampinae - Pedipedinae - Pythiinae - †Zaptychiinae
Genera Incertae sedis (14): †Juramarinula – †Leopoldium – †Melampoides – †Mesauriculstra – †Mesochilina – †Palaeoleuca – †Proauriculastra – †Proplecotrema – †Pythiopsis – †Rhytophorus – †Semiauricula – †Stolidoma – †Tortacella – †Traliopsis
Overview of genera (43)
Extant groups (25): Allochroa – Auriculastra – Blauneria – Carychiella – Carychium – Cassidula – Creedonia – Detracia – Ellobium – Koreozospeum – Laemodonta – Leuconopsis – Leucophytia – Marinula – Melampus – Microtralia – Myosotella – Ophicardelus – Ovatella – Pedipes – Pleuroloba – Pseudomelampus – Pythia – Tralia – Zospeum
Fossil groups (18): †Auriculinella – †Carychiopsis – †Juramarinula – †Leopoldium – †Melampoides – †Mesauriculstra – †Mesochilina – †Palaeoleuca – †Proauriculastra – †Proplecotrema – †Pythiopsis – †Rhytophorus – †Semiauricula – †Stolidoma – †Sulcomarinula – †Tortacella – †Traliopsis – †Zaptychius
Name
Ellobiidae Pfeiffer, 1854 (1822)
References
Pfeiffer, L. 1854. Synopsis Auriculaceorum. Malakozoologische Blätter 1: 146.
Bouchet, P. & Rocroi, J.-P. 2005. Classification and nomenclator of gastropod families. Malacologia 47(1–2): 1–397. BHL Reference page.
Frias Martins, A.M. de 1996: Anatomy and systematics of the western Atlantic Ellobiidae. Malacologia, 37(2): 163–332. BHL
Frias Martins, A.M. de 2007: Morphological and anatomical diversity within the Ellobiidae (Gastropoda, Pulmonata, Archaeopulmonata). Vita malacologica, (4): 1–28. abstract only seen
Links
Ellobiidae in the World Register of Marine Species
Vernacular names
日本語: オカミミガイ科
Nederlands: Oorslakken
Ellobiidae, common name the hollow-shelled snails, is a family of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Eupulmonata.[2] Ellobiidae is the only family in the superfamily Ellobioidea, according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[3]
Anatomy
In this family, the number of haploid chromosomes lies between 16 and 20 (according to the values in this table).[4]
They have a distinctive mode of locomotion arising due to a split sole. The front part of the sole adheres to the substrate and then the rear part of the sole is drawn up to the front part.[5]
Taxonomy
Species are traditionally classified into five taxonomic groups, the Pythiidae, the Laemodontidae, the Melampodidae, the Ellobiidae and the Carychiidae.[1] These taxa have been recognized either as families within Ellobioidea or as sub-families within the family Ellobiidae.[1]
The family Ellobiidae consists of the following subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):[3]
subfamily Ellobiinae Pfeiffer, 1854 (1822)
subfamily Carychiinae Jeffreys, 1830
subfamily Melampinae Stimpson, 1851 (1850)
subfamily Pedipedinae P. Fischer & Crosse, 1880
subfamily Pythiinae Odhner, 1925 (1880)
† subfamily Zaptychiinae Wenz, 1938
The high degree of homoplasy in morphological characters and frequent low variability has led to the description of approximately 800 species names available in the literature, of which 250 are likely to be valid.[1]
A molecular study by Dayrat at al. (2011)[6] for the Ellobioidea suggests a monophyletic origin of the entire group.[1]
Genera
Genera within the family Ellobiidae include:
subfamily Ellobiinae
Blauneria Shuttleworth, 1854
Ellobium Röding, 1798
subfamily Carychiinae
Carychium O. F. Müller, 1773
Zospeum Bourguignat, 1856
subfamily Melampinae
Melampus Montfort, 1810
subfamily Pedipedinae
Creedonia Martins, 1996
Leuconopsis Hutton, 1884
Marinula King & Broderip, 1832
Microtralia Dall, 1894
Pedipes Férussac, 1821
Pseudomelampus Pallary, 1900
Sarnia H. Adams & A. Adams, 1855 (temporary name)
subfamily Pythiinae Odhner, 1925
Allochroa Ancey, 1887
Auriculastra Martens, 1880
Cassidula Gray, 1847
Laemodonta Philippi, 1846
Myosotella Monterosato, 1906
Ophicardelus Beck, 1838
Ovatella Bivona, 1832
Pleuroloba Hyman, Rouse & Ponder, 2005
Pythia Röding, 1798 - type genus of the subfamily Pythiinae.
† subfamily Zaptychiinae
† Zaptychius Walcott, 1883 - type genus of the subfamily
subfamily ?
Auriculinella Tausch, 1886
Auriculodes Strand, 1928
Blauneria Shuttleworth, 1854
Detracia Gray in Turton, 1840
Tralia Gray, 1840
Genera brought into synonymy
Alexia Leach in Gray, 1847: synonym of Myosotella Monterosato, 1906
Auricula Lamarck, 1799: synonym of Ellobium Röding, 1798
Cremnobates Swainson, 1855: synonym of Marinula King & Broderip, 1832
Jaminia T. Brown, 1827: synonym of Myosotella Monterosato, 1906
Kochia Pallary, 1900: synonym of Myosotella Monterosato, 1906
Laimodonta H. Adams & A. Adams, 1855: synonym of Laemodonta Philippi, 1846
Leuconia Gray, 1840: synonym of Auriculinella Tausch, 1886
Leucopepla Peile, 1926: synonym of Auriculinella Tausch, 1886
Leucophytia Winckworth, 1949: synonym of Auriculinella Tausch, 1886
Monica H. Adams & A. Adams, 1855: synonym of Ovatella Bivona-Bernardi, 1832
Nealexia Wenz, 1920: synonym of Myosotella Monterosato, 1906
Phytia: synonym of Pythia Röding, 1798
Pira H. & A. Adams, 1855: synonym of Melampus Montfort, 1810
Plecotrema H. Adams & A. Adams, 1854: synonym of Laemodonta Philippi, 1846
Rangitotoa Powell, 1933: synonym of Microtralia Dall, 1894
Taxonomic note
Genus Sarnia H. Adams & A. Adams, 1855 (temporary name, Sarnia [H. & A. Adams, 1855 (September), The Genera of Recent Mollusca, 2: 239] is an incorrect subsequent spelling of Siona H. Adams & A. Adams, 1855 which is a junior homonym of Siona Duponchel, 1829 [Lepidoptera].)
Ecology
The taxon Ellobioidea comprises a group of morphologically and ecologically highly diverse snails, known to have successfully invaded the marine, brackish water and terrestrial habitats.[1]
These are mostly snails that live in salt marshes and similar maritime habitats, and thus have a tolerance for saline conditions.
References
This article incorporates CC-BY-2.0 text from the reference[1]
Weigand A. M., Jochum A., Slapnik R., Schnitzler J., Zarza E. & Klussmann-Kolb A. (2013). "Evolution of microgastropods (Ellobioidea, Carychiidae): integrating taxonomic, phylogenetic and evolutionary hypotheses". BMC Evolutionary Biology 13: 18. doi:10.1186/1471-2148-13-18.
Bouchet, P. (2012). Ellobiidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=198 on 2012-08-19
Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
Herbert,dai and, Kilburn, Dick (2004). Field Guide to the Land Snails and Slugs of Eastern South Africa. Scottsville, South Africa: Natal Museum. p. 98. ISBN 0-620-32415-5.
Dayrat B., Conrad M., Balayan S., White T. R., Albrecht C., Golding R., Gomes S., Harasewych M. G., de Frias Martins A. M. (2011). "Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling". Molecular Phylogenetics and Evolution 59: 425-437. doi:10.1016/j.ympev.2011.02.014.
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