Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Ringipleura
SuperOrdo: Nudipleura
Ordo: Nudibranchia
SubOrdo: Doridina
InfraOrdo: Doridoidei
Superfamilia: Doridoidea
Familiae (2): Discodorididae - Dorididae
Name
Doridoidea Rafinesque, 1815
Type Genus: Doris Linnaeus, 1758
References
Johnson, R.F. 2011: Breaking family ties: taxon sampling and molecular phylogeny of chromodorid nudibranchs (Mollusca, Gastropoda). Zoologica scripta, 40(2): 137–157. DOI: 10.1111/j.1463-6409.2010.00457.x
Valdés, Á. 2002. A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia). Zoological journal of the Linnean Society 136(4): 535–636. DOI: 10.1046/j.1096-3642.2002.00039.x Open access. Reference page.
Links
Doridoidea in the World Register of Marine Species
Doridoidea, commonly known as dorid nudibranchs (and previously known as the taxon Cryptobranchia), are a taxonomic superfamily of medium to large, shell-less sea slugs, marine gastropod mollusks in the clade Doridacea, included in the clade Nudibranchia.
Etymology
The word "Doridoidea" comes from the generic name Doris, which was in turn copied from the name of the sea nymph, Doris, in Greek mythology.
Families
Peltodoris nobilis from Yaquina Bay
According to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), families within the superfamily Doridoidea include:
Actinocyclidae O'Donoghue, 1929
Chromodorididae Bergh, 1891
Discodorididae Bergh, 1891
Dorididae Rafinesque, 1815
Changes later than 2005
The family Cadlinidae Bergh, 1891 was considered a synonym of the Chromodorididae. Research by R.F. Johnson in 2011[2] has shown that Cadlina does not belong to the family Chromodorididae. She has therefore brought back the name Cadlinidae from synonymy with Chromodorididae. The chromodorid nudibranchs without Cadlina are now monophyletic and turn out to be a possible sister to the Actinocyclidae
Currently recognized families
Actinocyclidae O'Donoghue, 1929
Cadlinidae Bergh, 1891
Chromodorididae Bergh, 1891
Discodorididae Bergh, 1891
Dorididae Rafinesque, 1815
Synonyms
Mating pair of cryptobranchs (Diaulula sandiegensis)
The next families are considered synonyms. But these names can still be found in many publications and on the internet.
Aldisidae Odhner, 1939 - synonym of Cadlinidae
Archidorididae Bergh, 1891 - synonym of Dorididae
Asteronotidae Thiele, 1931 - synonym of Discodorididae
Baptodorididae Odhner, 1926 - synonym of Discodorididae
Conualeviidae Collier & Farmer, 1964 - synonym of Dorididae
Geitodorididae Odhner, 1968 - synonym of Discodorididae
Halgerdidae Odhner, 1926 - synonym of Discodorididae
Homoiodorididae Bergh, 1882 -synonym of Dorididae and Dendrodorididae
Kentrodorididae Bergh, 1891 - synonym of Discodorididae
Platydorididae Bergh, 1891 - synonym of Discodorididae
Rostangidae Pruvot-Fol, 1951 - synonym of Discodorididae
Cryptobranch dorid nudibranchs
Cryptobranch dorid nudibranchs (previously known as the taxon Cryptobranchia), are nudibranch sea slugs within the clade Doridacea. These slugs are called "cryptobranch," meaning "hidden gill", because they are able to retract their gills into a gill pocket, in contrast to nudibranchs in the traditional group phanerobranchs (or Phanerobranchia), which taxon is probably paraphyletic (in other words, composed of more than one evolutionary lineage).[3]
A. Valdés distinguishes two major clades within the Cryptobranchia: the dorids that have no radula (the Porostomata); and those with a radula (the Labiostomata). The Labiostomata include the monophyletic families: Actinocyclidae, Chromodorididae, Dorididae and Discodorididae.[4]
The cryptobranchs include the following genera that are regarded as valid:
Doris Linnaeus, 1758
Asteronotus Ehrenberg, 1831
Atagema J. E. Gray, 1850
Jorunna Bergh, 1876
Discodoris Bergh, 1877
Platydoris Bergh, 1877
Thordisa Bergh, 1877
Diaulula Bergh, 1878
Aldisa Bergh, 1878
Rostanga Bergh, 1879*[clarification needed]
Aphelodoris Bergh, 1879
Halgerda Bergh, 1880
Peltodoris Bergh, 1880
Hoplodoris Bergh, 1880
Paradoris Bergh, 1884
Baptodoris Bergh, 1884
Geitodoris Bergh, 1891
Gargamella Bergh, 1894
Alloiodoris Bergh, 1904
Sclerodoris Eliot, 1904
Otinodoris White, 1948
Taringa Er. Marcus, 1955
Sebadoris Er. Marcus & Ev. Marcus, 1960
Conualevia Collier & Farmer, 1964
Thorybopus Bouchet, 1977
Goslineria Valdés, 2001
Pharodoris Valdés, 2001
Nophodoris Valdés & Gosliner, 2001
References
Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdes A. & Warén A. 2005. Classification and nomenclator of gastropod families. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. ISBN 3-925919-72-4. ISSN 0076-2997. 397 pp. http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278
Johnson R.F. (2011) Breaking family ties: taxon sampling and molecular phylogeny of chromodorid nudibranchs (Mollusca, Gastropoda). Zoologica Scripta 40(2): 137-157
Valdés, Angel (2002). "A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)". Zoological Journal of the Linnean Society. 136 (4). London: The Linnean Society of London: 535–536. doi:10.1046/j.1096-3642.2002.00039.x.
Valdés, A. (2002). "A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)". Zoological Journal of the Linnean Society. 136 (4). Blackwell Publishing: 535–536. doi:10.1046/j.1096-3642.2002.00039.x.
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