Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Caenogastropoda
Ordo: Neogastropoda
Superfamilia: Buccinoidea
Familia: Columbellidae
Subfamiliae (2): Atiliinae - Columbellinae
Genera Incertae sedis (34 + 3†): Alcira – Bathyglypta – Bifurcium – Cilara – Clathranachis – Clavistrombina – Costoanachis – Eurypyrene – Euspiralta – Exaesopus – Glyptanachis – Graphicomassa – Indomitrella – Ithiaesopus – Lavesopus – Metanachis – Minipyrene – Mitropsis – Nodochila – Pardalinops – Parvanachis – Pictocolumbella – Pleurifera – Psarostola – Pyreneola – Ramoliva – Ruthia – Sincola – Steironepion – Strombina – Sulcomitrella – Suturoglypta – Zanassarina – Zetekia – †Antizafra – †Columbellisipho – †Trahaldia
Overview of genera (73 + 3†)
Aesopus – Alcira – Alia – Amphissa – Anachis – Antimitrella – Aoteatilia – Aoteatilia – Astyris – Bathyglypta – Bifurcium – Cilara – Clathranachis – Clavistrombina – Columbella – Conella – Cosmioconcha – Costoanachis – Cotonopsis – Decipifus – Euplica – Eurypyrene – Euspiralta – Exaesopus – Exomilopsis – Gatliffena – Glyptanachis – Graphicomassa – Indomitrella – Ithiaesopus – Lavesopus – Liratilia – Macrozafra – Mazatlania – Metanachis – Metulella – Microcithara – Minipyrene – Mitrella – Mitropsis – Mokumea – Nassarina – Nitidella – Nodochila – Parametaria – Pardalinops – Parvanachis – Parviterebra – Paxula – Pictocolumbella – Pleurifera – Psarostola – Pseudamycla – Pseudanachis – Pyrene – Pyreneola – Ramoliva – Retizafra – Rhombinella – Ruthia – Salitra – Seminella – Sincola – Steironepion – Strombina – Sulcomitrella – Suturoglypta – Zafra – Zafrona – Zanassarina – Zella – Zemitrella – Zetekia – †Antizafra – †Columbellisipho – †Trahaldia
[Source: WoRMS 7-27-2019 for complete genera list -- (subfamilia allocation is from prior Wikispecies investigation)]
Name
Columbellidae Swainson, 1840
References
Swainson, W.J. 1840. A treatise on Malacology; or, the natural classification of shells and shell fish. London, Longman 1–419. DOI: 10.5962/bhl.title.8027 BHL Reference page.
Chen, C., Watanabe, H.K. & Araya, J.F. 2017. First columbellid species (Gastropoda: Buccinoidea) from deep-sea hydrothermal vents, discovered in Okinawa Trough, Japan. Zootaxa 4363(4): 592–596. DOI: 10.11646/zootaxa.4363.4.13 Full text PDF from ResearchGate. Reference page.
Links
Columbellidae in the World Register of Marine Species
The Columbellidae, the dove snails or dove shells, are a family of minute to small sea snails, marine gastropod mollusks in the order Neogastropoda.[2] [3]
Pardalinops testudinaria
These snails are characterized by their elongated, spiral shells and their carnivorous feeding habits. The family Columbellidae comprises numerous genera and species, many of which share similar shell morphologies and ecological niches.
Distribution
Species in this family are found worldwide,[4] but are most abundant in shallow tropical water.
Subfamilies
According to the taxonomy of the Gastropoda by Bouchet & Rocroi (2005), the family Columbellidae consists of two subfamilies:
Atiliinae Cossmann, 1901 - synonyms: Pyrenidae Suter, 1909; Anachidae Golikov & Starobogatov, 1972
Columbellinae Swainson, 1840
Columbellidae incertae sedis (temporary name)
Columbellidae incertae sedis mariato (Pilsbry & H. N. Lowe, 1932)
Genera
The family Columbellidae contains these genera:
subfamily Atiliinae
subfamily Columbellinae
† Ademitrella Ludbrook, 1941
Aesopus Gould, 1860
Alcira H. Adams, 1861
Alia H. Adams & A. Adams, 1853
Amphissa H. Adams & A. Adams, 1853
Anachis H. Adams & A. Adams, 1853
Antimitrella Powell, 1937
† Antizafra H. J. Finlay, 1926
Aoteatilia Powell, 1939
Ascalista Drivas & Jay, 1990
Astyris H. Adams and A. Adams, 1853
† Auingeria Harzhauser & Landau, 2021
Bathyglypta Pelorce, 2017
† Bellacolumbella Harzhauser & Landau, 2021
Bifurcium P. Fischer, 1884
† Cigclirina Woodring, 1928
Cilara Thiele, 1924
Clathranachis Kuroda & Habe, 1954
Clavistrombina Jung, 1989
† Clinurella Sacco, 1890
Columbella Lamarck, 1799 - type genus
† Columbellisipho Cossmann, 1889
Conella Swainson, 1840
Cosmioconcha Dall, 1913
Costoanachis Sacco, 1890
Cotonopsis Olsson, 1942
Decipifus Olsson and McGinty, 1958
† Defensina Harzhauser & Landau, 2021
Euplica Dall, 1889
Eurypyrene Woodring, 1928
Euspiralta K. Monsecour & Pelorce, 2013
Exaesopus de Maintenon, 2019
Exomilopsis Powell, 1964
Falsuszafrona Pelorce, 2020
Gatliffena Iredale, 1929
Glyptanachis Pilsbry & Lowe, 1932
Graphicomassa Iredale, 1929
Indomitrella Oostingh, 1940
Ithiaesopus Olsson & Harbison, 1953
† Juliamitrella L. D. Campbell, 1993
Liratilia Finlay, 1927
Macrozafra Finlay, 1927
Maintenonia K. Monsecour & D. Monsecour, 2024
† Martaia Harzhauser & Landau, 2021
Mazatlania Dall, 1913
Metanachis Thiele, 1924
Metulella Gabb, 1873
Microcithara P. Fischer, 1884
Minimanachis Pelorce, 2020
Mitrella Risso, 1826
Mitropsis Pease, 1868
Mokumea Habe, 1991
Nassarina Dall, 1889
Nitidella Swainson, 1840
Nodochila Rehder, 1980
† Orthurella Sacco, 1890
Parametaria Dall, 1916
Pardalinops De Maintenon, 2008
Parvanachis Radwin, 1968
Parviterebra Pilsbry, 1904
Paxula Finlay, 1927
Pictocolumbella Habe, 1945
Pleurifera Drivas & Jay, 1997
Psarostola Rehder, 1943: synonym of Steironepion Pilsbry & H. N. Lowe, 1932
Pseudamycla Pace, 1902
Pyrene Röding, 1798
Pyreneola Iredale, 1918
Radwinia Shasky, 1970
Ramoliva Cotton & Godfrey, 1932
Redfernella Espinosa & Ortea, 2020
Retizafra Hedley, 1913
Rhombinella Radwin, 1968
Ruthia Shasky, 1970
Salitra Marincovich, 1973
† Scabrella Sacco, 1890
Seminella Pease, 1867
Sincola Olsson & Harbison, 1953
Smithena Boyer, Pelorce & Gori, 2022
Speirazafra K. Monsecour & Raines, 2024
Steironepion Pilsbry and Lowe, 1932
Strombina Mörch, 1852
† Strombinella Dall, 1896
Sulcomitrella Kuroda, Habe & Oyama, 1971
Suturoglypta Radwin, 1968
† Thiarella Sacco, 1890
† Trahaldia Pacaud, 2017
† Turricolumbus Cossmann, 1901
Turrijaumelia Sarasúa, 1975
Urpiella Arias Ávila, 2021
Zafra A. Adams, 1860
Zafrona Iredale, 1916
Zanassarina Pilsbry & H. N. Lowe, 1932
Zella Iredale, 1924
Zemitrella Finlay, 1927
Zetekia Dall, 1918
Synonyms
Atilia H. Adams & A. Adams, 1853 - type genus of the subfamily Atiliinae: synonym of Anachis H. Adams & A. Adams, 1853
Columbellopsis Bucquoy, Dautzenberg and Dollfus, 1882: synonym of Mitrella Risso, 1826
Minipyrene Coomans, 1967: synonym of Mitromorpha P. P. Carpenter, 1865
References
Swainson (1840). Treat. Malacol. 152, 313.
Bouchet, P.; Gofas, S. (2010). Columbellidae. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=23009 on 2010-11-04
Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdes A. & Warén A. 2005. Classification and nomenclator of gastropod families. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. ISBN 3-925919-72-4. ISSN 0076-2997. 397 pp. http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278
"Archived copy". Archived from the original on 2007-12-22. Retrieved 2007-12-25.
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