Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Euteleosteomorpha
Subcohors: Neoteleostei
Infracohors: Eurypterygia
Sectio: Ctenosquamata
Subsectio: Acanthomorphata
Divisio/Superordo: Acanthopterygii
Ordo: Syngnathiformes
Subordo: Mulloidei
Familia: Mullidae
Genus: Upeneus
Species (39):
U. arge – U. asymmetricus – U. australiae – U. crosnieri – U. davidaromi – U. doriae – U. farnis – U. filifer – U. flavolineatus – U. francisi – U. guttatus – U. heemstra – U. indicus – U. itoui – U. japonicus – U. lombok – U. luzonius – U. margarethae – U. mascareinsis – U. moluccensis – U. nigromarginatus – U. parvus – U. pori – U. quadrilineatus – U. randalli – U. saiab – U. seychellensis – U. spottocaudalis – U. stenopsis – U. suahelicus – U. subvittatus – U. sulphureus – U. sundaicus – U. supravittatus – U. taeniopterus – U. tragula – U. vanuatu – U. vittatus – U. xanthogrammus
Name
Upeneus Cuvier, 1829
Type species: Mullus vittatus Forsskål, 1775
Synonyms
Hypeneus Agassiz, 1846
Megalepis Bianconi, 1857
Pennon Whitley, 1941
Upeneoides Bleeker, 1849
References
Bos, A.R., 2014: Upeneus nigromarginatus, a new species of goatfish (Perciformes: Mullidae) from the Philippines. Raffles Bulletin of Zoology 62: 745–753. Full article: [1].
Cuvier, G. (1829): Le Règne Animal, distribué d'après son organisation, pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée. Edition 2. Vol. 2: i-xv + 1-406.
Uiblein, F.; Causse, R. 2013: A new deep-water goatfish of the genus Upeneus (Mullidae) from Vanuatu, South Pacific. Zootaxa 3666(3): 337–344. DOI: 10.11646/zootaxa.3666.3.4 Reference page.
Uiblein, F., Gledhill, D.C. & Peristiwady, T. 2017. Two new goatfishes of the genus Upeneus (Mullidae) from Australia and Indonesia. Zootaxa 4318(2): 295–311. DOI: 10.11646/zootaxa.4318.2.4. Full article (PDF). Reference page.
Uiblein, F. and P.C. Heemstra, 2010: A taxonomic review of the Western Indian goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin 11: 35–71.
Uiblein, F. & P.C. Heemstra, 2011: Description of a new goatfish species, Upeneus randalli n. sp. (Mullidae), from the Persian Gulf, with remarks and identification keys for the genus Upeneus. Scientia Marina 75(3): 583–594. Full article: [2].
Uiblein, F. & P.C. Heemstra, 2011: A new goatfish, Upeneus seychellensis n. sp. (Mullidae), from the Seychelles Bank, with remarks on Upeneus guttatus and a key to Western Indian Ocean Upeneus species. Mar. Biol. Res. 7(7): in press.
Uiblein, F. & Lisher, M. 2013: A new goatfish of the genus Upeneus (Mullidae) from Angoche, northern Mozambique. Zootaxa 3717(1): 85–95. DOI: 10.11646/zootaxa.3717.1.7 Reference page.
Uiblein, F.; McGrouther, M. 2012: A new deep-water goatfish of the genus Upeneus (Mullidae) from northern Australia and the Philippines, with a taxonomic account of U. subvittatus and remarks on U. mascareinsis. Zootaxa 3550: 61–70. Preview Reference page.
Uiblein, F. & White, W.T. 2015: A new goatfish of the genus Upeneus (Mullidae) from Lombok, Indonesia and first verified record of U. asymmetricus for the Indian Ocean. Zootaxa 3980(1): 51–66. DOI: 10.11646/zootaxa.3980.1.3. Preview (PDF) Full article (PDF) Reference page.
Links
Upeneus – Taxon details on Integrated Taxonomic Information System (ITIS).
Upeneus species list in FishBase,
Froese, R. & Pauly, D. (eds.) 2021. FishBase. World Wide Web electronic publication, www.fishbase.org, version 02/2021.
Vernacular names
Upeneus is a genus of goatfishes native to the Atlantic, Indian and Pacific oceans.
Species
There are currently 43 recognized species in this genus:[2][3]
Upeneus asymmetricus Lachner, 1954 (Asymmetrical goatfish)
Upeneus australiae B. J. Kim & Nakaya, 2002 (Australian goatfish)
Upeneus caudofasciatus Uiblein & Gledhill, 2019
Upeneus davidaromi Golani, 2001 (Darom's goatfish)
Upeneus doriae Günther, 1869 (Gilded goatfish)
Upeneus farnis Uiblein & Peristiwady, 2017 (Farnis' goatfish)[4]
Upeneus filifer J. D. Ogilby, 1910 (Pennant goatfish)
Upeneus francisi J. E. Randall & Guézé, 1992 (Francis' goatfish)
Upeneus gubal Uiblein, 2019
Upeneus guttatus F. Day, 1868 (Two-tone goatfish)
Upeneus heemstra Uiblein & Gouws, 2014[5] (Heemstra goatfish)
Upeneus heterospinus Uiblein & Pavlov, 2019
Upeneus indicus Uiblein & Heemstra, 2010 (Tall-fin goatfish)
Upeneus itoui M. Yamashita, Golani & Motomura, 2011[6] (Oriental goatfish)
Upeneus japonicus |Houttuyn, 1782 (Japanese goatfish)
Upeneus lombok Uiblein & W. T. White, 2015[7] (Lombok goatfish)
Upeneus luzonius D. S. Jordan & Seale, 1907 (Dark-barred goatfish)
Upeneus margarethae Uiblein & Heemstra, 2010 (Margaretha's goatfish)
Upeneus mascareinsis Fourmanoir & Guézé, 1967 (Mascarene goatfish)
Upeneus moluccensis Bleeker, 1855 (Goldband goatfish)
Upeneus mouthami J. E. Randall & Kulbicki, 2006 (Moutham's goatfish)
Upeneus niebuhri Guézé, 1967[5] (Niebuhr's goatfish)
Upeneus nigromarginatus A. R. Bos, 2014[8] (Black-margined goatfish)
Upeneus oligospilus Lachner, 1954 (Short-fin goatfish)
Upeneus parvus Poey, 1852 (Dwarf goatfish)
Upeneus pori Ben-Tuvia & Golani, 1989 (Por's goatfish)
Upeneus quadrilineatus C. T. Cheng & C. X. Wang, 1963 (Four-stripe goatfish)
Upeneus randalli Uiblein & Heemstra, 2011
Upeneus saiab Uiblein & Lisher, 2013[9] (Saiab goatfish)
Upeneus seychellensis Uiblein & Heemstra, 2011 (Tailstripe goatfish)
Upeneus spottocaudalis Uiblein & Danel Gledhill, 2017 (Tailspot goatfish)[4]
Upeneus stenopsis Uiblein & McGrouther, 2012 (|Narrowtail goatfish)
Upeneus suahelicus Uiblein & Heemstra, 2010 (Swahili goatfish)
Upeneus subvittatus Temminck & Schlegel, 1843 (Deep-water goatfish)
Upeneus sulphureus G. Cuvier, 1829 (Sulphur goatfish)
Upeneus sundaicus Bleeker, 1855 (Ochrebanded goatfish)
Upeneus supravittatus Uiblein & Heemstra, 2010 (Long-fin goatfish)
Upeneus taeniopterus G. Cuvier, 1829 (Finstripe goatfish)
Upeneus torres Uiblein & Gledhill, 2015[10] (Torres goatfish)
Upeneus tragula J. Richardson, 1846 (Freckled goatfish)
Upeneus vanuatu Uiblein & Causse, 2013[11] (Vanuatu goatfish)
Upeneus vittatus Forsskål, 1775 (Yellowstriped goatfish)
Upeneus xanthogrammus C. H. Gilbert, 1892
References
Cuvier, G. (1829): Le Règne Animal, distribué d'après son organisation, pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée. Edition 2. 2: i-xv + 1-406.
Froese, Rainer and Pauly, Daniel, eds. (2019). Species of Upeneus in FishBase. December 2019 version.
Uiblein, Franz; Gledhill, Daniel C.; Pavlov, Dimitri A.; Hoang, Tuan Anh; Shaheen, Shaker (2019-10-08). "Three new goatfishes of the genus Upeneus (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in U. margarethae". Zootaxa. 4683 (2): 151–196. doi:10.11646/zootaxa.4683.2.1. ISSN 1175-5334.
Franz Uiblein; Daniel Gledhill & Teguh Peristiwady (2017). "Two new goatfishes of the genus Upeneus (Mullidae) from Australia and Indonesia". Zootaxa. 4318 (2): 295–311. doi:10.11646/zootaxa.4318.2.4.
Uiblein, F. & Gouws, G. (2014): A new goatfish species of the genus Upeneus (Mullidae) based on molecular and morphological screening and subsequent taxonomic analysis. Marine Biology Research, 10 (7): 655-681.
Yamashita, M., Golani, D. & Motomura, H. (2011). "A new species of Upeneus (Perciformes: Mullidae) from southern Japan". Zootaxa. 3107: 47–58. doi:10.11646/zootaxa.3107.1.3.
Uiblein, F. & White, W.T. (2015). "A new goatfish of the genus Upeneus (Mullidae) from Lombok, Indonesia and first verified record of U. asymmetricus for the Indian Ocean" (PDF). Zootaxa. 3980 (1): 51–66. doi:10.11646/zootaxa.3980.1.3. hdl:11250/2356213. PMID 26249938.
Bos, A.R. (2014). "Upeneus nigromarginatus, a new species of goatfish (Perciformes: Mullidae) from the Philippines" (PDF). Raffles Bulletin of Zoology. 62: 745–753. Archived from the original (PDF) on 2014-12-13. Retrieved 2014-12-13.
Uiblein, F. & Lisher, M. (2013): A new goatfish of the genus Upeneus (Mullidae) from Angoche, northern Mozambique. Zootaxa, 3717 (1): 85–95.
Uiblein F. & Gledhill D.C. (2015). "A new goatfish of the genus Upeneus (Mullidae) from Australia and Vanuatu, with inter- and intraspecific comparisons". Marine Biology Research. 11 (5): 475–491. doi:10.1080/17451000.2014.958088. S2CID 84788326.
Uiblein, F. & Causse, R. (2013): A new deep-water goatfish of the genus Upeneus (Mullidae) from Vanuatu, South Pacific. Zootaxa, 3666 (3): 337–344.
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