Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Euteleosteomorpha
Subcohors: Neoteleostei
Infracohors: Eurypterygia
Sectio: Ctenosquamata
Subsectio: Acanthomorphata
Divisio/Superordo: Acanthopterygii
Subdivisio: Percomorphaceae
Series: Gobiaria
Ordo: Gobiiformes
Subordo: Gobioidei
Familia: Gobiidae
Subfamilia: Gobiinae
Genus: Trimma
Species: T. abyssum – T. agrena – T. anaima – T. annosum – T. anthrenum – T. aturirii – T. avidori – T. barralli – T. bathum – T. benjamini – T. bisella – T. blematium – T. burridgeae – T. caesiura – T. cana – T. caudipunctatum – T. caudomaculatum – T. cheni – T. chledophilum – T. corallinum – T. corerefum – T. dalerocheila – T. emeryi – T. erdmanni – T. eviotops – T. fangi – T. fasciatum – T. filamentosum – T. finistrinum – T. fishelsoni – T. flammeum – T. flavatrum – T. flavicaudatum – T. fraena – T. fucatum – T. gigantum – T. grammistes – T. griffithsi – T. habrum – T. haima – T. haimassum – T. halonevum – T. hamartium – T. hayashii – T. hoesei – T. hollemani – T. hotsarihiensis – T. imaii – T. insularum – T. kardium – T. kitrinum – T. kudoi – T. lantana – T. macrophthalmum – T. maiandros – T. marinae – T. matsunoi – T. meityae – T. mendelssohni – T. meristum – T. milta – T. multiclitellum – T. nasa – T. nauagium – T. naudei – T. necopina – T. nomurai – T. okinawae – T. omanensis – T. papayum – T. pentherum – T. pharus – T. preclarum – T. quadrimaculatum – T. randalli – T. readerae – T. rubromaculatum – T. sanguinellus – T. sheppardi – T. sostra – T. squamicana – T. stobbsi – T. striatum – T. tauroculum – T. taylori – T. tevegae – T. trioculatum – T. unisquamis – T. volcana – T. winchi – T. winterbottomi – T. woutsi – T. xanthochrum – T. xanthum – T. yanagitai – T. yanoi
Add (1): T. irinae
Name
Trimma Jordan & Seale, 1906
Type species: Trimma caesiura (neuter) Jordan & Seale, 1906. Type by original designation (also monotypic).
References
Hoese, D.F., Bogorodsky, S.V. & Mal, A.O., 2015: Description of a new species of Trimma (Perciformes: Gobiidae) from the Red Sea, with a discussion of the generic separation of Trimma and Priolepis, with discussion of sensory papillae terminology. Zootaxa 4027(4): 538–550. DOI: 10.11646/zootaxa.4027.4.4. Preview (PDF) Reference page.
Hoese, D.F.; Winterbottom, R.; Reader, S. 2011: Trimma maiandros, a new species of pygmy goby (Gobiidae) from the Indo-west Pacific. Aqua, 17(2): 103–110.
Jordan, D.S. & A. Seale 1906: The fishes of Samoa. Description of the species found in the archipelago, with a provisional check-list of the fishes of Oceania. Bulletin of the Bureau of Fisheries, 25(for 1905): 173–455 + index 457-488, Pls. 33-53.
Suzuki, T. & H. Senou 2009: Two new species of the genus Trimma (Perciformes: Gobiidae) from Japan and Palau. Bulletin of the Kanagawa Prefectural Museum, Natural Science, 38: 107–112.
ITIS
Winterbottom, R. 2016. Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa 4144(1): 1–53. DOI: 10.11646/zootaxa.4144.1.1. Reference page.
Winterbottom, R. 2017. Two new species of Trimma (Pisces; Gobiidae) from Fiji, south-western Pacific Ocean. Zootaxa 4269(4): 559–570. DOI: 10.11646/zootaxa.4269.4.9. Reference page.
Winterbottom, R. 2018. A new species of Trimma (Pisces; Gobiidae) from the South-West Islands of Palau, western Pacific Ocean. Zootaxa 4370(2): 123–136. DOI: 10.11646/zootaxa.4370.2.2 Reference page.
Winterbottom, R.; Erdmann, M.V.; Cahyani, N.K.D. 2014: Trimma helenae (Pisces; Gobioidei), a new species of gobiid fish from Indonesia. Zootaxa 3760(3): 420–428. DOI: 10.11646/zootaxa.3760.3.8 Reference page.
Winterbottom, R. et al. 2014: A cornucopia of cryptic species - a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys 381: 79-111. DOI: 10.3897/zookeys.381.6445 Reference page.
Winterbottom, R.; Erdmann, M.V.; Cahyani, N.K.D. 2014: Three new species of Trimma (Pisces; Gobioidei) from Indonesia. Zootaxa 3838(3): 367–384. DOI: 10.11646/zootaxa.3838.3.7 Reference page.
Winterbottom, R. & Erdmann, M.V. 2018. Two new species of blue-eyed Trimma (Pisces; Gobiidae) from New Guinea. Zootaxa 4444(4): 471–483. DOI: 10.11646/zootaxa.4444.4.7 Paywall Reference page.
Winterbottom, R., Erdmann, M.V. & Cahyani, N.K.D. 2015: New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature. Zootaxa 3973(2): 201–226. DOI: 10.11646/zootaxa.3973.2.1. Preview (PDF) Reference page.
Winterbottom, R. & Hoese, D. 2015: A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species. Zootaxa 3934(1): 1–102. DOI: 10.11646/zootaxa.3934.1.1. Reference page.
Vernacular names
English: Pygmy Gobies
Trimma is a genus of fish in the family Gobiidae native to the Indian and Pacific Ocean. Together with members of the genus Eviota, they are known commonly as pygmygobies or dwarfgobies.
Species
Trimma caudomaculatum
Trimma habrum
Trimma lantana
Trimma nasa
Trimma naudei female 23.3 mm, Nha Trang, Vietnam
Trimma okinawae
Trimma tevegae
There are currently 98 recognized species in this genus:
Trimma abyssum G. R. Allen, 2015 (Deep-reef pygmygoby) [1]
Trimma agrena R. Winterbottom & I. S. Chen, 2004 (Fishnet pygmygoby)
Trimma anaima R. Winterbottom, 2000 (Sharp-eye pygmygoby) [2]
Trimma annosum R. Winterbottom, 2003 (Grey-bearded pygmygoby) [2]
Trimma anthrenum R. Winterbottom, 2006 (Honey-bee pygmygoby)
Trimma aturirii R. Winterbottom, Erdmann & Cahyani, 2015 (Aturiri's pygmygoby) [3]
Trimma avidori (Goren, 1978)
Trimma barralli R. Winterbottom, 1995
Trimma benjamini R. Winterbottom, 1996 (Ring-eye pygmygoby) [2]
Trimma bisella R. Winterbottom, 2000
Trimma burridgeae R. Winterbottom, 2016 [4]
Trimma caesiura D. S. Jordan & Seale, 1906 (Caesiura pygmygoby) [2]
Trimma cana R. Winterbottom, 2004 (Candycane pygmygoby)
Trimma capostriatum (Goren, 1981) (Spotted red-lined pygmygoby) [2]
Trimma caudipunctatum T. Suzuki & Senou, 2009
Trimma caudomaculatum Yoshino & Araga, 1975 [4]
Trimma cheni R. Winterbottom, 2011 (Face-stripe pygmygoby)
Trimma chledophilum G. R. Allen, 2015 (Mud pygmygoby) [1]
Trimma corallinum (J. L. B. Smith, 1959) (Polka-dot pygmygoby)
Trimma corerefum R. Winterbottom, 2016 [4]
Trimma dalerocheila R. Winterbottom, 1984
Trimma emeryi R. Winterbottom, 1985 (Emery's pygmygoby) [2]
Trimma erdmanni R. Winterbottom, 2011 (Erdmann's pygmygoby)
Trimma erwani Viviani, J. T. Williams & Planes, 2016 (Slanted pygmygoby) [5]
Trimma fangi R. Winterbottom & I. S. Chen, 2004 (Spot-faced pygmygoby) [2]
Trimma fasciatum T. Suzuki, Sakaue & Senou, 2012 [6]
Trimma filamentosus R. Winterbottom, 1995
Trimma fishelsoni Goren, 1985
Trimma flammeum (J. L. B. Smith, 1959)
Trimma flavatrum K. Hagiwara & R. Winterbottom, 2007 (Wasp pygmygoby) [2]
Trimma flavicaudatum (Goren, 1982)
Trimma fraena R. Winterbottom, 1984
Trimma fucatum R. Winterbottom & Southcott, 2007 (Harlot pygmygoby)
Trimma gigantum R. Winterbottom & Zur, 2007 (Giant pygmygoby)
Trimma grammistes (Tomiyama, 1936)
Trimma griffithsi R. Winterbottom, 1984 (Griffiths' pygmygoby)
Trimma habrum R. Winterbottom, 2011 (Delicate pygmygoby)
Trimma haima R. Winterbottom, 1984
Trimma haimassum R. Winterbottom, 2011 (Blood-spot pygmygoby)
Trimma halonevum R. Winterbottom, 2000 (Red-spot pygmygoby)
Trimma hayashii K. Hagiwara & R. Winterbottom, 2007 (Four-eye pygmygoby)
Trimma helenae R. Winterbottom, Erdmann & Cahyani, 2014 (Helen's pygmygoby) [7]
Trimma hoesei R. Winterbottom, 1984 (Fork-tail pygmygoby) [2]
Trimma hollemani R. Winterbottom, 2016 [4]
Trimma hotsarihiensis R. Winterbottom, 2009 (Helen Reef pygmygoby)
Trimma imaii T. Suzuki & Senou, 2009 (Imai's pygmygoby)
Trimma insularum R. Winterbottom & Hoese, 2015 (Cocos pygmygoby) [2]
Trimma irinae R. Winterbottom, 2014 (Irina's pygmygoby) [8]
Trimma kardium R. Winterbottom, Erdmann & Cahyani, 2015 (Heart pygmygoby) [3]
Trimma kitrinum R. Winterbottom & Hoese, 2015 (Citron pygmygoby) [2]
Trimma kudoi T. Suzuki & Senou, 2008
Trimma lantana R. Winterbottom & C. A. Villa, 2003 (Lantana pygmygoby) [2]
Trimma lutea Viviani, J. T. Williams & Planes, 2016 (Yellow-barred pygmygoby) [5]
Trimma macrophthalmus (Tomiyama, 1936) (Large-eye pygmygoby) [2]
Trimma maiandros Hoese, R. Winterbottom & Reader, 2011 (Zigzag pygmygoby) [2]
Trimma marinae R. Winterbottom, 2005 (Princess pygmygoby)
Trimma matsunoi T. Suzuki, Sakaue & Senou, 2012 [6]
Trimma mendelssohni (Goren, 1978)
Trimma meranyx R. Winterbottom, Erdmann & Cahyani, 2014 (Day-night pygmygoby) [9]
Trimma meristum R. Winterbottom & Hoese, 2015 (Split-ray pygmygoby) [2]
Trimma milta R. Winterbottom, 2002 (Red-earth pygmygoby) [2]
Trimma multiclitellum G. R. Allen, 2015 (Multi-saddle pygmygoby) [1]
Trimma nasa R. Winterbottom, 2005 (Nasal-bar pygmygoby) [2]
Trimma nauagium G. R. Allen, 2015 (Shipwreck pygmygoby) [1]
Trimma naudei J. L. B. Smith, 1957 (Naude's pygmygoby)
Trimma necopinum (Whitley, 1959) (Australian pygmygoby) [2]
Trimma nomurai T. Suzuki & Senou, 2007 (Lilac pygmygoby) [2]
Trimma okinawae (Aoyagi, 1949) (Orange-red pygmygoby) [2]
Trimma omanensis R. Winterbottom, 2000
Trimma pajama R. Winterbottom, Erdmann & Cahyani, 2014 (Pajama pygmygoby) [9]
Trimma papayum R. Winterbottom, 2011 (Pawpaw pygmygoby)
Trimma pentherum R. Winterbottom & Hoese, 2015 (Mourning pygmygoby) [2]
Trimma preclarum R. Winterbottom, 2006 (Exquisite pygmygoby) [2]
Trimma quadrimaculatum Hoese, Bogorodsky & A. O. Mal, 2015 (Four-spotted pygmygoby) [10]
Trimma randalli R. Winterbottom & Zur, 2007 (Randall's pygmygoby)
Trimma readerae R. Winterbottom & Hoese, 2015 [2]
Trimma rubromaculatum G. R. Allen & Munday, 1995 (Red-spotted dwarfgoby)
Trimma sanguinellus R. Winterbottom & Southcott, 2007 (Sanguinello pygmygoby)
Trimma sheppardi R. Winterbottom, 1984 (Sheppard's pygmygoby)
Trimma sostra R. Winterbottom, 2004 (Sostra pygmygoby)
Trimma squamicana R. Winterbottom, 2004 (Candy-scale pygmygoby)
Trimma stobbsi R. Winterbottom, 2001 (Stobb's pygmygoby) [2]
Trimma striatum (Herre, 1945) (Red-lined pygmygoby) [2]
Trimma tauroculum R. Winterbottom & Zur, 2007 (Bulls-eye pygmygoby)
Trimma taylori Lobel, 1979 (Yellow pygmygoby) [2]
Trimma tevegae Cohen & W. P. Davis, 1969 (Blue-stripe pygmygoby) [4]
Trimma trioculatum R. Winterbottom, Erdmann & Cahyani, 2015 (Three-eyed pygmygoby) [3]
Trimma unisquame (Gosline, 1959) (Black-margin pygmygoby) [2]
Trimma volcana R. Winterbottom, 2003 (Volcano pygmygoby)
Trimma winchi R. Winterbottom, 1984
Trimma winterbottomi J. E. Randall & Downing, 1994 (Winterbottom's pygmygoby)
Trimma woutsi R. Winterbottom, 2002
Trimma xanthochrum R. Winterbottom, 2011 (Step-spot pygmygoby)
Trimma xanthum R. Winterbottom & Hoese, 2015 (Yellow-red pygmygoby) [2]
Trimma yanagitai T. Suzuki & Senou, 2007
Trimma yanoi T. Suzuki & Senou, 2008 (Yano's pygmygoby)
Trimma yoshinoi T. Suzuki, Ko. Yano & Senou, 2015 [11]
Trimma zurae R. Winterbottom, Erdmann & Cahyani, 2014 (Marg's pygmygoby) [9]
References
Wikimedia Commons has media related to Trimma.
Allen, G.R. (2015): Four new species of Trimma (Pisces: Gobiidae) from Papua New Guinea and Indonesia. Journal of the Ocean Science Foundation, 14: 28-48.
Winterbottom, R. & Hoese, D.F. (2015): A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species. Zootaxa, 3934 (1): 1–102.
Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2015): New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature. Zootaxa, 3973 (2): 201–226.
Winterbottom, R. (2016): Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa, 4144 (1): 1-53.
Viviani, J., Williams, J.T. & Planes, S. (2016): Two new pygmygobies (Percomorpha: Gobiidae: Trimma) from French Polynesia. Journal of the Ocean Science Foundation, 23: 1–11.
Suzuki, T., Sakaue, J. & Senou, H. (2012): Two New Species of the Gobiid Fish Genus Trimma (Actinopterygii: Perciformes: Gobiidae) from Japan and Palau. Bulletin of the National Museum of Nature and Science, Ser. A, Supplement No. 6: 67-77.
Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2014): Trimma helenae (Pisces; Gobioidei), a new species of gobiid fish from Indonesia. Zootaxa, 3760 (3): 420–428.
Winterbottom, R. (2014): Trimma irinae, (Pisces; Gobioidei) a new species of gobiid fish from Milne Bay Province, Papua New Guinea. Zootaxa, 3802 (2): 209–216.
Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2014): Three new species of Trimma (Pisces; Gobioidei) from Indonesia. Zootaxa, 3838 (3): 367–384.
Hoese, D.F., Bogorodsky, S.V. & Mal, A.O. (2015): Description of a new species of Trimma (Perciformes: Gobiidae) from the Red Sea, with a discussion of the generic separation of Trimma and Priolepis, with discussion of sensory papillae terminology. Zootaxa, 4027 (4): 538–550.
Suzuki, T., Yano, K. & Senou, H. (2015): Trimma yoshinoi, a new gobiid fish from Japan (Perciformes: Gobiidae). Journal of the Ocean Science Foundation, 14: 66-73.
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