Thymallus thymallus, Photo: Christian Maier
Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Euteleosteomorpha
Subcohors/Superordo: Protacanthopterygii
Ordo: Salmoniformes
Familia: Salmonidae
Subfamilia: Thymallinae
Genus: Thymallus
Species: Thymallus thymallus
Name
Thymallus thymallus (Linnaeus, 1758)
Original combination: Salmo thymallus
References
Linnaeus, C. 1758. Systema Naturae per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Tomus I. Editio decima, reformata. Holmiæ: impensis direct. Laurentii Salvii. i–ii, 1–824 pp DOI: 10.5962/bhl.title.542: 311. Reference page.
Thymallus thymallus in FishBase,
Froese, R. & Pauly, D. (eds.) 2022. FishBase. World Wide Web electronic publication, www.fishbase.org, version 08/2021.
Vernacular names
Alemannisch: Äsche
беларуская: Ліпень
български: Липан
bosanski: Lipljen
català: Thymallus thymallus
čeština: Lipan podhorní
dansk: Stalling
Deutsch: Europäische Äsche
English: Grayling
español: Tímalo
eesti: Harjus, Ärn, Harjakas, Õrn, Hõbeforell
suomi: Harjus
français: Ombre commun
hrvatski: Lipljen
magyar: Pénzes pér
italiano: Temolo
lietuvių: Kiršlys
Nederlands: Vlagzalm
norsk nynorsk: Harr
norsk: Harr
polski: Lipień pospolity
русский: Европейский хариус
slovenčina: Lipeň tymiánový
slovenščina: Lipan
српски / srpski: Липљен
svenska: Harr
українська: Харіус європейський
Thymallus thymallus, the grayling or European grayling,[3] is a species of freshwater fish in the salmon family Salmonidae. It is the only species of the genus Thymallus (the graylings) native to Europe, where it is widespread from the United Kingdom and France to the Ural Mountains in Russia, and Balkans on the south-east, but does not occur in the southern parts of the continent. It was introduced to Morocco in 1948, but it does not appear to have become established there.
Description
The grayling grows to a maximum recorded length of 60 cm (24 in) and a maximum recorded weight of 6.7 kg (15 lb). Of typical Thymallus appearance, the grayling proper is distinguished from the similar Arctic grayling (T. arcticus arcticus) by the presence of 5–8 dorsal and 3–4 anal spines, which are absent in the other species; T. thymallus also has a smaller number of soft rays in these fins. Individuals of the species have been recorded as reaching an age of 14 years.
The grayling prefers cold, clean, running riverine waters, but also occurs in lakes and, exceptionally, in brackish waters around the Baltic Sea. Omnivorous, the fish feeds on vegetable matter, as well as crustaceans, insects and spiders, mollusks, zooplankton, and smaller fishes, such as Eurasian minnows. Grayling are also prey for larger fish, including the huchen (Hucho hucho).
With the Arctic grayling, T. thymallus is one of the economically important Thymallus species, being raised commercially and fished for sport.
The grayling is a protected species listed in Appendix III of the Bern Convention. It has become critically endangered in the Baltic Sea.[4]
Adult grayling
The term "grayling" is often used to refer generically to the Thymallus species, and T. thymallus is sometimes called the European grayling for clarity. There are many obsolete synonyms for the species.
The generic name Thymallus derives from the Greek θύμαλλος, "thyme smell", a name derived from the fragrance of wild thyme that freshly caught graylings are believed to smell similar to. Thymallus thymallus is the type species of its genus.
Angling
The grayling is known as the 'lady of the stream'.[5] In Europe, they used to be persecuted by anglers for the false perception that they stopped trout colonizing stretches of rivers and streams. However, research has shown that grayling and trout feed on different prey items and generally prefer different microhabitats within rivers and streams but do occupy similar niches to smaller, less-predatory trout.[6][7]
In England and Wales, they can be fished for throughout the coarse fishing season (16 June to 14 March), providing thrilling sport on the fly when the trout season is closed. There is no closed season for grayling in Scotland; where they have been introduced. There are no grayling in Ireland. Well-known grayling flies include the grayling witch, klinkhamers, various nymphs and 'red tags', along with other trout patterns. Flies tied to resemble small pink shrimps have also been found to be useful. A method known as 'Czech-nymphing' has been known to be helpful to anglers where grayling shoal up in colder periods. The method involves moving a series of Czech nymphs under the tip of the fly rod with the flow of the river and the nymphs should entice the grayling to take one. Fly-anglers may wade in the river to perform this method where they can access deeper water. Wading does not necessarily spook the grayling as they are generally less cautious than trout and are not as easily put off by human presence.
In France, too, the season is limited depending upon several factors. The Allier is one of the rare places in Southern Europe where the common grayling occurs in a natural habitat.
References
Freyhof, J. (2011). "Thymallus thymallus". IUCN Red List of Threatened Species. 2011: e.T21875A9333742. doi:10.2305/IUCN.UK.2008.RLTS.T21875A9333742.en.
Linnaeus, Carl, Systema Naturae … , 10th ed. (Stockholm (Holmia), Sweden: Lars Salvius (Laurentius Salvius), 1758), vol. 1, p. 311.
Smoliński, Szymon; Glazaczow, Adam (4 December 2019). "Cascading effects of temperature alterations on trophic ecology of European grayling (Thymallus thymallus)". Scientific Reports. 9 (1): 18358. Bibcode:2019NatSR...918358S. doi:10.1038/s41598-019-55000-5. ISSN 2045-2322. PMC 6892815. PMID 31798001.
HELCOM (2013). "HELCOM Red List of Baltic Sea species in danger of becoming extinct" (PDF). Baltic Sea Environmental Proceedings (140): 72. Archived from the original (PDF) on 2014-10-07. Retrieved 2014-07-30.
"Publications - The Grayling Society - Angling - Conservation - Research". Retrieved 15 April 2017.
"The Grayling Research Trust: Competition". Grayling Research Trust. Archived from the original on 22 February 2014.
"Food‐ and habitat‐segregation in sympatric grayling and brown trout". Retrieved 15 April 2017.
Further reading
"Grayling" . Encyclopædia Britannica. Vol. 11 (9th ed.). 1880. p. 78.
"Grayling" . Encyclopædia Britannica. Vol. 12 (11th ed.). 1911. p. 395.
World Conservation Monitoring Centre (1996). "Thymallus thymallus". IUCN Red List of Threatened Species. 1996. Retrieved 5 May 2006.
Froese, Rainer; Pauly, Daniel (eds.) (2004). "Thymallus thymallus" in FishBase. October 2004 version.
"Thymallus thymallus". Integrated Taxonomic Information System. Retrieved 11 December 2004.
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