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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Otomorpha
Subcohors: Ostariophysi
Sectio: Otophysa
Ordo: Cypriniformes
Subordo: Cobitoidei

Familia: Nemacheilidae
Genus: Barbatula
Species (37): B. altayensis – B. altayensis – B. araxensis – B. barbatula – B. bergamensis – B. bergiana – B. brandtii – B. cinica – B. compressirostris – B. dgebuadzei – B. emuensis – B. erdali – B. euphratica – B. farsica – B. frenata – B. germencica – B. gibba – B. golubtsovi – B. karabanowi – B. kermanshahensis – B. kosswigi – B. liaoyangensis – B. linjiangensis – B. mediterraneus – B. nuda – B. panthera – B. paucilepis – B. persa – B. phoxinoides – B. potaninorum – B. quignardi – B. restricta – B. sawadai – B. seyhanensis – B. sturanyi – B. toni – B. tschaiyssuensis – B. zetensis – B. zhangwuensis
Name

Barbatula Linck, 1790
References
Primary references

Linck, H.F. 1790. Versuch einer Eintheilung der Fische nach den Zähnen. Magazin für das Neueste aus der Physik und Naturgeschichte Gotha 6(3): 28–38. Google Books Open access. Reference page.

Additional references

Chen, H., Zhang, H., Chen, Y-X. & Freyhof, J. 2019. A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species. Zootaxa 4565(1): 1–36. DOI: 10.11646/zootaxa.4565.1.1 Paywall Reference page.
Cao, L.; Causse, R.; Zhang, E. 2012: Revision of the loach species Barbatula nuda (Bleeker 1865) (Pisces: Balitoridae) from North China, with a description of a new species from Inner Mongolia. Zootaxa 3586: 236–248. Preview PDF Reference page.
Prokofiev, A.M. 2018. A new species of Barbatula from the headwaters of the Bulgan River in western Mongolia (Teleostei: Nemacheilidae). Zootaxa 4407(2): 267–274. DOI: 10.11646/zootaxa.4407.2.7 Reference page.

Links

Barbatula species list in FishBase,
Froese, R. & Pauly, D. (eds.) 2024. FishBase. World Wide Web electronic publication, www.fishbase.org, version 02/2024.
Barbatula and its species (including synonyms) in Catalog of Fishes, Eschmeyer, W.N., Fricke, R. & van der Laan, R. (eds.) 2024. Catalog of Fishes electronic version.
Barbatula – Taxon details on Nomenclator Zoologicus (NZ).

Vernacular names
eesti: Trulling

Barbatula is a genus of fish in the family Nemacheilidae native to Europe and Asia.[2][3] They are found in streams, rivers and lakes, and the genus also includes Europe's only cavefish, which only was discovered in the Danube–Aachtopf system in Germany in 2015.[4][5]

Barbatula formerly included many more species, but these have been moved to other genera, notably Oxynoemacheilus.[2]
Species

There are currently 18 recognized species in this genus:[3]

Barbatula altayensis S. Q. Zhu, 1992
Barbatula barbatula (Linnaeus, 1758)
Barbatula bergamensis Erk'Akan, Nalbant & Özeren, 2007
Barbatula compressirostris (Warpachowski, 1897)
Barbatula dgebuadzei (Prokofiev, 2003)
Barbatula farsica (Nalbant & Bianco, 1998)
Barbatula gibba Cao, Causse & Zhang, 2012
Barbatula golubtsovi (Prokofiev, 2003)
Barbatula markakulensis (Men'shikov, 1939)
Barbatula nuda (Bleeker, 1864)
Barbatula potaninorum (Prokofiev, 2007)
Barbatula quignardi (Băcescu-Meşter, 1967)
Barbatula restricta Prokofiev, 2015[6]
Barbatula sawadai (Prokofiev, 2007)
Barbatula sturanyi (Steindachner, 1892)
Barbatula tomiana (Ruzsky (ru), 1920)
Barbatula toni (Dybowski, 1869)[7]
Barbatula zetensis (Šorić, 2000)

References

Bailly, Nicolas (2014). "Barbatula Linck, 1790". WoRMS. World Register of Marine Species. Retrieved 29 January 2024.
Kottelat, M. (2012): Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). Archived 2016-05-09 at the Wayback Machine Raffles Bulletin of Zoology, Suppl. No. 26: 1-199.
Froese, Rainer and Pauly, Daniel, eds. (2024). Species of Barbatula in FishBase. February 2024 version.
Behrmann-Godel, J.; A.W. Nolte; J. Kreiselmaier; R. Berka; J. Freyhof (2017). "The first European cave fish". Current Biology. 27 (7): R257–R258. doi:10.1016/j.cub.2017.02.048. PMID 28376329.
Andy Coghlan (3 April 2017). "First ever cavefish discovered in Europe evolved super-fast". New Scientist. Retrieved 17 April 2017.
Prokofiev, A. M. (2015). "A new species of Barbatula from the Russian Altai (Teleostei: Nemacheilidae)" (PDF). Zootaxa. 4052 (4): 457. doi:10.11646/zootaxa.4052.4.3.
Prokofiev, A. M. (2016). "Redescription and systematic position of nominal loach species Nemacheilus compressirostris and N. sibiricus (Nemacheilidae)". Journal of Ichthyology. 56 (4): 488–497. doi:10.1134/S0032945216040111.

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