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Miacidae

Superregnum: Eukaryota
Cladus: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Mammalia
Subclassis: Theria
Infraclassis: Placentalia
Ordo: Carnivora
Subordo: †Miacoidea
Familia: Miacidae
Genera: Chailicyon - Eosictis - Ictognathus - Messelogale - Miacis - Miocyon - Oödectes - Palaearctonyx - Paramiacis - Paroodectes - Procynodictis - Prodaphaenus - Quercygale - Tapocyon - Uintacyon - Vassacyon - Vulpavus - Xinyuictis - Ziphacodon

Name

Miacidae Cope, 1880

Miacids (Miacidae) were primitive carnivores which lived during the late Paleocene and Eocene epochs about 62—33 million years ago. Miacids existed for approximately 29 million years.

Miacids are thought to have evolved into today's modern carnivorous mammals of the order Carnivora. They were small carnivores, superficially marten-like or civet-like with long, little bodies and long tails. Some species were arboreal while others lived on the ground.

They probably fed on invertebrates, lizards, birds, and smaller mammals like shrews and opossums. Their teeth and skulls show that the miacids were less developed than modern carnivorans. They had carnivoran-type carnassials but lacked fully ossified auditory bullae (rounded protrusions).

Classification

Miacidae as traditionally conceived is not a monophyletic group; it is a paraphyletic array of stem taxa. Traditionally, Miacidae and Viverravidae had been classified in a superfamily, Miacoidea. Today, Carnivora and Miacoidea are grouped together in the crown-clade Carnivoramorpha, and the Miacoidea are regarded as basal carnivoramorphs. Some species of the genus Miacis are closely related to the order Carnivora, but only the species Miacis cognitus is a true carnivoran, as it is classified in the Caniformia.

The transition from miacids to Carnivora was a general trend in the middle and late Eocene with taxa from both North America and Eurasia involved. The divergence of carnivorans from miacids is now inferred to have occurred in the middle-Eocene (ca. 42 million years ago). Traditionally the Viverravidae (viverravids) had been thought to be the earliest carnivorans with fossil records first appearing in the Paleocene of North America about 60 million years ago, but recent cranial morphology evidence now places them outside the order Carnivora.[1]

Taxonomy

Family Miacidae†

Genus Eosictis
Genus Messelogale[2]
Genus Miacis
Genus Miocyon
Genus Oodectes
Genus Palaearctonyx
Genus Paramiacis
Genus Paroodectes
Genus Procynodictis[3]
Genus Prodaphaenus
Genus Quercygale[4]
Genus Tapocyon
Genus Uintacyon
Genus Vassacyon
Genus Vulpavus
Genus Xinyuictis
Genus Ziphacodon

References

^ Polly, David, Gina D. Wesley-Hunt, Ronald E. Heinrich, Graham Davis and Peter Houde (2006). "Earliest Known Carnivoran Auditory Bulla and Support for a Recent Origin of Crown-Clade Carnivora (Eutheria, Mammalia)". Palaeontology 49 (5): 1019–1027. doi:10.1111/j.1475-4983.2006.00586.x.
^ Morlo, M.; Schaal, S.; Mayr, G.; Seiffert, C. (2004). "An annotated taxonomic list of the Middle Eocene (MP11) Vertebrata of Messel". Courier Forschungsinstitut Senckenberg 252: 95–108. Retrieved 2008-05-18.
^ Wesley, G.D.; Flynn, J.J. (2003). "A revision Of Tapocyon (Carnivoramorpha), including analysis of the first cranial specimens and identification of a new species". Journal of Paleontology 77 (4): 769–783. doi:10.1666/0022-3360(2003)077<0769:AROTCI>2.0.CO;2.
^ Wesley-Hunt, G.D.; Werdelin, L. (2005). "Basicranial morphology and phylogenetic position of the upper Eocene carnivoramorphan Quercygale". Acta Palaeontologica Polonica 50 (4): 837–846. Retrieved 2008-05-18.

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