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Penelope jacquacu

Penelope jacquacu, Photo: Raimond Spekking (*)

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Galliformes
Subordo: Craci

Familia: Cracidae
Genus: Penelope
Species: Penelope jacquacu
Subspecies: P. j. granti – P. j. jacquacu – P. j. orienticola – P. j. speciosa
Name

Penelope jacquacu Spix, 1825
References

Spix, J.B. von 1825. Avium species novae, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis. Suscepto. Collegit et descripsit Dr. J. B. de Spix. Tomus 2. 85 pp. + 115 tt. Illustrations: Matthias Schmidt. Ed. Typis Franc. Seraph. Hübschmanni. Monachii. DOI: 10.5962/bhl.title.63182 p. 52 BHL Reference page. pl. 68 BHL

Vernacular names
English: Spix's Guan
español: Pava amazónica
português: Jacu-de-spix

Spix's guan (Penelope jacquacu) is a species of bird in the family Cracidae. It is "the prototypical cracid of the Amazonian lowlands."[2] The common name commemorates the German naturalist Johann Baptist von Spix (1782-1826).[3]
Taxonomy and systematics

Four subspecies of Spix's guan are recognized, "Grant's" guan (P. j. granti) and the "Spix's guan" group of P. j. jacquacu, P. j. orienticola, and P. j. speciosa.[2] It was at times considered conspecific with dusky-legged guan (P. obscura) or crested guan (P. purpurascens). P. j. speciosa was once considered a race of dusky-legged guan, and "Grant's guan" was formerly considered a separate species.[2][4]
Description

The four subspecies of Spix's guan differ in weight and plumage, and there are intergrades between them. Their length is 66–76 cm (26–30 in). The male and female nominate P. j. jacquacu weigh 1.24–1.36 kg (2.7–3.0 lb) and 1.14 kg (2.5 lb) respectively. The male P. j. orienticola weighs 1.40–1.78 kg (3.1–3.9 lb) and the female 1.27–1.72 kg (2.8–3.8 lb). The nominate subspecies is bronzy olive green above and bright reddish below. "Grant's" is much darker; the upper parts have a blue-green gloss and the red underside is dark. P. j. speciosa's plumage is similar to that of the nominate subspecies and P. j. orienticola's size and color scheme are between those of the nominate and "Grant's".[2]
Distribution and habitat

Spix's guan is found in the upper Orinoco River and western Amazon River basins of Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, Suriname, and Venezuela. From north to south the subspecies sequence is granti, orienticola, jacquacu, and speciosa.[2][4] It inhabits both terra firma and várzea (seasonally flooded) forest. In the upper elevations of Peru and Bolivia it is in cloud forest and at lower elevations it is in humid rainforest. The species is assumed to be sedentary.[2]
Behavior and ecology
Breeding

The timing of Spix's guan's breeding varies geographically; examples include from January to May in Venezuela and August and September in Peru. The one nest that has been described was made of leaves and placed about 5 m (16 ft) up in a tree. The incubation and fledging periods are not known.[2]
Diet and feeding

Spix's guan forages singly or in small family groups, mostly from the middle to the tops of trees but in some regions on the ground. Its diet is mainly fruits and seeds though it has also been reported to follow army ants to take arthropods.[2]
Status

The IUCN has rated Spix's guan as being of Least Concern. Its estimated global range spans approximately 5.9 million km2 (2.3 million mi2) and it is common in at least parts of this range. However, the total population is unknown and is thought to be decreasing. The species is widely hunted for food.[1]
References
Wikimedia Commons has media related to Penelope jacquacu.

BirdLife International (2016). "Penelope jacquacu". IUCN Red List of Threatened Species. 2016: e.T22678386A92771552. doi:10.2305/IUCN.UK.2016-3.RLTS.T22678386A92771552.en. Retrieved 16 November 2021.
del Hoyo, J. and G. M. Kirwan (2020). Spix's Guan (Penelope jacquacu), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.spigua1.01
Beolens, Bo; Watkins, Michael (2003). Whose Bird? Men and Women Commemorated in the Common Names of Birds. London: Christopher Helm. p. 319.
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 19 January 2021. A classification of the bird species of South America. American Ornithological Society. http://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved January 19, 2021

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