Heliodoxa imperatrix

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Superordo: Caprimulgimorphae
Ordo: Apodiformes

Familia: Trochilidae
Subfamilia: Trochilinae
Genus: Heliodoxa
Species: Heliodoxa imperatrix
Name

Heliodoxa imperatrix (Gould, 1856)

Type locality: Andean forests in the neighborhood of Quito, Ecuador.

Synonyms

Eugenia imperatrix (protonym)

References

Gould, J. 1855. On a new Genus and Species of Trochilidae from Ecuador. Proceedings of the Zoological Society of London Pt 23 no. 296: p. 192 BHL Reference page.

Vernacular names
English: Empress Brilliant
español: Brillante emperador

The empress brilliant (Heliodoxa imperatrix) is a species of hummingbird in the "brilliants", tribe Heliantheini in subfamily Lesbiinae. It is found in Colombia and Ecuador.[3][4]
Taxonomy and systematics

In the early part of the 20th century the empress brilliant was treated as the only member of genus Eugenia, which was later merged into the present Heliodoxa.[5] The species is monotypic.[3]
Description

Male empress brilliants are 15 to 17 cm (5.9 to 6.7 in) long and weigh about 9.3 g (0.33 oz). Females are 12 to 13.5 cm (4.7 to 5.3 in) long and weigh about 8.3 g (0.29 oz). The adult male's forehead, face, and breast are mostly glittering dark green, with a square glittering pale violet patch in the center of the throat. Its crown and nape are dark green and the rest of the upperparts dark bronzy green. Its belly is glittering golden green. The central tail feathers are dark bronze and the rest longer and black with a bronze gloss. The adult female's upperparts are bronzy green. The center of its throat and breast are grayish with much bronzy green flecking, the flanks solid bronzy green, and the belly golden green. The central tail feathers are bronzy green and the others dull black with a bronze gloss.[6]

Immature males have a dull dark bronzy green head, throat, and breast; bright buff chin and sides of the throat; and a duller and bronzier green belly than the adult. Immature females also have a buffy chin, and the green feathers of the breast are duller than the adult's and have buffy fringes.[6]
Distribution and habitat

The empress brilliant is found on the Pacific slope of the Andes from Colombia's Chocó Department south into Ecuador as far as Pichincha Province. It inhabits a variety of very wet landscapes including foothill forest, the interior and edges of cloudforest, and mature secondary forest. In elevation it ranges from 400 to 2,000 m (1,300 to 6,600 ft).[6]
Behavior
Movement

The movements of the empress brilliant, if any, have not been described.[6]
Feeding

Both sexes of empress brilliant forage from the mid-level to the forest canopy, and females also forage in the understorey. A primary source of nectar are the flowers of Marcgravia and Marcgraviastrum vines, where it perches to feed. It also feeds by hovering under drooping flowers of Ericaceae. In addition to feeding on nectar, it also captures small insects by gleaning from foliage and hawking from a perch.[6]
Breeding

The empress brilliant's breeding season spans from January to April. The few known nests were cups of tree fern scales and other plant material held together with spider silk and lined with soft seed fluff. Very little else is known about the species' breeding phenology.[6]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to empress brilliant on xeno-canto
Vocalization

One empress brilliant vocalization is a "repeated single 'tsit' note". It is also given singly while hovering and feeding.[6]
Status

The IUCN has assessed the empress brilliant as being of Least Concern. However, it has a restricted range and its population size and trend are unknown. No immediate threats have been identified.[1] It appears to be reasonably common and occurs in some protected areas. "[Its] very wet forest habitat still mostly intact".[6]
References

BirdLife International (2016). "Empress Brilliant Heliodoxa imperatrix". IUCN Red List of Threatened Species. 2016: e.T22687715A93165555. doi:10.2305/IUCN.UK.2016-3.RLTS.T22687715A93165555.en. Retrieved 7 May 2022.
"Appendices | CITES". cites.org. Retrieved 2022-01-14.
Gill, F.; Donsker, D.; Rasmussen, P., eds. (January 2022). "Hummingbirds". IOC World Bird List. v 12.1. Retrieved January 15, 2022.
HBW and BirdLife International (2020) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip [.xls zipped 1 MB] retrieved 27 May 2021
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 January 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved February 1, 2022
Stiles, F.G. and P. F. D. Boesman (2020). Empress Brilliant (Heliodoxa imperatrix), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.empbri1.01 retrieved 7 May 2022