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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Superordo: Caprimulgimorphae
Ordo: Apodiformes

Familia: Trochilidae
Subfamilia: Trochilinae
Genus: Heliangelus
Species: Heliangelus exortis
Name

Heliangelus exortis, (Fraser, 1840)

Type locality: Guaduas, Colombia.

Synonymy

Trochilus exortis (protonym)
Ornismya parzudaki Lesson, 1840 Rev.Zool. p. 72 BHL
Heliotrypha parzudaki (Lesson, 1840)
Heliangelus exortis soderstromi Oberholser, 1902

References

Fraser, L. 1840. Characters of New Species of Humming-Birds. Proceedings of the Zoological Society of London Pt.8 no.86: 14–19. BHL Reference page. p. 14

Vernacular names
English: Tourmaline Sunangel
español: Colibrí turmalina

The tourmaline sunangel (Heliangelus exortis) is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Colombia and Ecuador.[3][4]
Taxonomy and systematics

The tourmaline sunangel is monotypic.[3] However, at times it has been considered conspecific with the flame-throated sunangel (H. micraster).[5]
Description

The tourmaline sunangel is 10 to 11 cm (3.9 to 4.3 in) long. Males weigh 3.5 to 8.5 g (0.12 to 0.30 oz) and females 4 to 5.2 g (0.14 to 0.18 oz). Their bill is blackish, straight, and of medium length. Both sexes have dark shining green upperparts and dusky gray underparts with round green spots. Their tails' central feathers are dark bronze and the others blackish. Adult males have a glittering green frontlet just above the bill, a dark purple-blue chin, and a glittering rosy pink gorget with a glittering emerald green border below it. Adult females lack the green frontlet, their chin is blackish, and their throat whitish with green to dusky speckles and sometimes a few glittering rosy feathers. Juveniles are like the adult female but the male has a white chin.[5]
Distribution and habitat

The tourmaline sunangel is found in all three Andes ranges of Colombia and south on the eastern slope to Ecuador's Morona-Santiago Province. It inhabits the interior and edges of humid to wet premontane forest, especially cloudforest and elfin forest. It also occurs in bushy clearings and shrubby pastures. In elevation it mainly ranges between 2,300 and 3,400 m (7,500 and 11,200 ft) but is found as low as 1,500 m (4,900 ft).[5]
Behavior
Movement

The tourmaline sunangel is almost entirely sendentary but in Colombia makes occasional movements into the tropical zone.[5]
Feeding

The tourmaline sunangel is aggressively territorial at feeding locations. It takes nectar from a variety of flowering plants, usually those low to the ground both in semi-open landscapes and within the forest. It usually clings to flowers to feed rather than hovering at them. It captures insects by hawking from a perch and by gleaning from vegetation.[5]
Breeding

The tourmaline sunangel's breeding season appears to span from March to August. The clutch of two white eggs is incubated by the female. Nothing else is known about the species' breeding phenology.[5]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to tourmaline sunangel on xeno-canto
Vocalization

A vocalization believed to be the tourmaline sunangel's song is "a repeated series of single notes...typically a few high-pitched upslurred notes followed by a slightly lower-pitched downslurred note, 'suwee....suwee..tsew...suwee...tsew...'." Its call is "a short dry gravelly trill 'trrr', repeated at intervals" and it also makes a continuous dry chatter.[5]
Status

The IUCN has assessed the tourmaline sunangel as being of Least Concern. Its population size is not known but is believed to be stable.[1] It is locally fairly common and occurs in several protected areas.[5]
References

BirdLife International (2017). "Tourmaline Sunangel Heliangelus exortis". IUCN Red List of Threatened Species. 2017: e.T22687893A112391164. doi:10.2305/IUCN.UK.2017-1.RLTS.T22687893A112391164.en. Retrieved 13 November 2021.
"Appendices | CITES". cites.org. Retrieved 2022-01-14.
Gill, F.; Donsker, D.; Rasmussen, P. (July 2021). "IOC World Bird List (v 12.1)". doi:10.14344/IOC.ML.11.2. Retrieved January 15, 2022.
HBW and BirdLife International (2020) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip [.xls zipped 1 MB] retrieved May 27, 2021
Heynen, I. and P. F. D. Boesman (2020). Tourmaline Sunangel (Heliangelus exortis), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.tousun1.01 retrieved January 21, 2022

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