Heliangelus clarisse

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Superordo: Caprimulgimorphae
Ordo: Apodiformes

Familia: Trochilidae
Subfamilia: Trochilinae
Genus: Heliangelus
Species: Heliangelus clarisse
Subspecies: H. c. clarisse – H. c. verdiscutus – H. c. violiceps
Name

Heliangelus clarisse (Longuemare, 1841)

Type locality: Bogotá, Colombia.

Synonyms

Ornismya clarisse (protonym)
Heliangelus claudia Hartert, 1895 Novit.Zool p. 484 BHL

References

Longuemare, A.F.G. 1841. Oiseau mouche nouveau. Revue Zoologique par La Société Cuvierienne 4: p. 306 BHL Reference page.

Vernacular names
English: Longuemare's Sunangel
español: Colibrí de Clarissa

Longuemare's sunangel (Heliangelus clarisse) is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Colombia and Venezuela.[3][4]
Taxonomy and systematics

The taxonomy of Longuemare's sunangel is not settled. The International Ornithological Committee (IOC), the Clements taxonomy, and BirdLife International's Handbook of the Birds of the World (HBW) treat it as a species and assign two subspecies to it, the nominate H. c. clarisse and H. c. violiceps. The South American Classification Committee (SACC) of the American Ornithological Society treats them as subspecies of the amethyst-throated sunangel (H. amethysticollis). The IOC and HBW also recognize H. c. verdiscutus as a subspecies, while Clements doesn't.[3][5][4][6]
Description

Longuemare's sunangel is 9 to 10 cm (3.5 to 3.9 in) long and weighs 5.0 to 6.0 g (0.18 to 0.21 oz). It has a short (less than 2 cm (0.79 in)), straight, blackish bill. The adult male of the nominate H. c. clarisse has a narrow glittering blue-green frontlet just above the bill, a dusky crown, and dark bronzy green upperparts. Its throat and upper breast gorget is glittering rosy and has a white pectoral band below it. The rest of the underparts are dusky buff with round green spots. The inner tail feathers are dark bronzy green and the outer ones blackish and occasionally have pale tips. The female has dull brown throat feathers with rufous or bronze green inclusions. Its belly has less spotting than the male's. Juveniles are plumaged like the adult female.[7][8]

The male H. c. violiceps has a strongly blue frontlet and a purple gorget. The female's frontlet is dull; its dull brown throat patch is smaller than that of the nominate and a band of dark green separates it from the white pectoral band. The male H. c. verdiscutus has an emerald green frontlet, a deep bronzy purple crown, and a dark line along the upper edge of the gorget. The female has the same dark line and a smaller gorget than the nominate's with a dark green border.[7]
Distribution and habitat

The nominate Longuemare's sunangel H. c. clarisse is found in Colombia's Eastern Andes and adjacent western Venezuela. H. c. violiceps is found further north, in Serranía del Perijá that defines the Colombia-Venezuela border. H. c. verdiscutus is restricted to the Tamá Massif, an extension of Colombia's eastern Andes on the border with Venezuela.[3][7]

Longuemare's sunangel primarily inhabits the edges of humid cloudforest and elfin forest. It can also be found in more open landscapes including fields with scattered bushes and trees, open woodland, overgrown ravines, and páramo. In elevation it ranges from 1,800 to 3,600 m (5,900 to 11,800 ft).[7][9]
Behavior
Movement

Longuemares' sunangel is sedentary.[7]
Feeding

Longuemare's sunangel feeds on nectar and also includes insects in its diet. It defends feeding territories that are dense with nectar sources. It typically feeds at heights up to 6 m (20 ft), often along streams. It captures insects by hawking from a perch.[7]
Breeding

The breeding season of Longuemare's sunangel extends at least from May to August. Its nest of moss and fine plant material is usually hung below a leaf or other structure that provides shelter from the weather. The clutch of two eggs is incubated by the female. The incubation period and time to fledging are not known.[7]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to Longuemare's sunangel on xeno-canto
Vocalization

Longuemare's sunangel's call is "a repeated dry, upward-inflected 'tsik' or 'tsit'."[7]
Status

The IUCN has assessed Longuemare's sunangel as being of Least Concern. Its population size is not known but is believed to be stable.[1] It is considered fairly common to common and no potential threats are known.[7]
References

BirdLife International (2016). "Longuemare's Sunangel Heliangelus clarisse". IUCN Red List of Threatened Species. 2016: e.T60859989A95164006. doi:10.2305/IUCN.UK.2016-3.RLTS.T60859989A95164006.en. Retrieved 20 January 2022.
"Appendices | CITES". cites.org. Retrieved 2022-01-14.
Gill, F.; Donsker, D.; Rasmussen, P., eds. (August 2022). "Hummingbirds". IOC World Bird List. v 12.2. Retrieved August 9, 2022.
HBW and BirdLife International (2020) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip [.xls zipped 1 MB] retrieved May 27, 2021
Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022
del Hoyo, J., I. Heynen, N. Collar, P. F. D. Boesman, G. M. Kirwan, and C. J. Sharpe (2020). Amethyst-throated Sunangel (Heliangelus amethysticollis), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.amtsun1.01 retrieved January 20, 2022
Hilty, Steven L. (2003), Birds of Venezuela, Princeton University Press, p. 426, ISBN 0-691-09250-8
Hilty, Steven L.; Brown, William L. (1986), A Guide to the Birds of Colombia, Princeton University Press, p. 66, ISBN 0-691-08371-1, retrieved 2007-10-06