Heliangelus amethysticollis

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Superordo: Caprimulgimorphae
Ordo: Apodiformes

Familia: Trochilidae
Subfamilia: Trochilinae
Genus: Heliangelus
Species: Heliangelus amethysticollis
Subspecies H. a. amethysticollis – H. a. decolor – H. a. laticlavius – H. a. violiceps
Name

Heliangelus amethysticollis (d’Orbigny & Lafresnaye, 1838)

Type locality: Yuracares, Cochabamba, Bolivia.

Synonyms

Ornismya amethysticollis (protonym)

References

d’Orbigny, A. & de Lafresnaye, F. 1838. Synopsis Avium, in ejus per Americam meridionalem itinere, collectarum et ab ipso viatore necnon. Magasin de zoologie 8(2):1–34, pl. 77-79. BHL Reference page. p. 31 BHL

Vernacular names
English: Amethyst-throated Sunangel
español: Colibrí gorjiamatista
français: Héliange à gorge améthyste

The amethyst-throated sunangel (Heliangelus amethysticollis) is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Bolivia, Ecuador, and Peru.[3][4]
Taxonomy and systematics

The amethyst-throated sunangel's taxonomy is not settled. The International Ornithological Committee (IOC), BirdLife International's Handbook of the Birds of the World (HBW), and the Clements taxonomy assign these four subspecies to it:[3][4]

H. a. laticlavius, Salvin (1891)
H. a. decolor, Zimmer, JT (1951)
H. a. apurimacensis, Weller (2009)
H. a. amethysticollis, d'Orbigny & Lafresnaye (1838)

The South American Classification Committee (SACC) of the American Ornithological Society includes three other subspecies. They are H. a. violiceps and H. a. clarisse that IOC, HBW, and the Clements taxonomy treat as Longuemare's sunangel (H. clarisse) and H. a. spencei that they treat as the Mérida sunangel (H. spencei).[5][3][4][6]
Peru, 2013
Description

The amethyst-throated sunangel, as defined by the IOC, is about 10 cm (3.9 in) long and weighs 3.8 to 7.7 g (0.13 to 0.27 oz). The species has a straight, rather short, black bill. The adult male of nominate subspecies H. a. amethysticollis has shiny dark green upperparts with a bright green frontlet just above the bill. Its throat and upper breast gorget is rosy amethyst. That area has a narrow black border on the neck and a broader cinnamon band below it. The lower breast is glittery green and the rest of the underparts buff with green discs. The central tail feathers are bronze-green and the outer ones blue-black. The adult female is similar to the male but has a rusty throat with black flecks. Immatures are similar to the female.[7]

Subspecies H. a. laticlavius has a reddish purple gorget, a whitish breastband, and a buffy grayish belly. H. a. decolor has a paler frontlet than the nominate; its gorget is a deeper violet, its breast band whiter, and its belly grayer. H. a. apurimacensis has a turquoise-green frontlet, a whitish buff breast, and a pale buff belly.[7]
Distribution and habitat

The four subspecies of amethyst-throated sunangel are found thus:[3][7]

H. a. laticlavius, the Andes of southern Ecuador south to northern Department of Cajamarca in Peru
H. a. decolor, the eastern slope of the Andes in central Peru from south of the Marañón River to the Department of Junín
H. a. apurimacensis, the valleys of the Apurímac River and upper Urubamba River in the departments of Ayacucho and Cusco, southeastern Peru
H. a. amethysticollis, the eastern slope of the Andes from southern Peru into northwestern Bolivia as far as Cochabamba Department

The species inhabits wet and humid montane forest including cloudforest and elfin forest. Though it is found at forest edges and open bushy terrain, it prefers the interior of closed forest. In elevation it mostly ranges between 1,800 and 3,200 m (5,900 and 10,500 ft) but has been often recorded as high as 3,700 m (12,100 ft) in Peru.[7]
Behavior
Movement

The amethyst-throated sunangel is sedentary.[7]
Feeding

The amethyst-throated sunangel feeds on nectar and also includes a high percentage of insects in its diet. It defends feeding territories that are dense with nectar sources; with its short bill it primarily feeds at flowers with shallow corollas. It typically feeds at heights up to 6 m (20 ft) and always near cover at the forest edge or interior. It captures insects by hawking from a perch.[7]
Breeding

The amethyst-throated sunangel nests in Peru during February and March. It builds a nest of fine plant fiber, moss, and lichens attached to a strand of moss dangling from a branch. It typically places it about 1.5 m (5 ft) above ground and under a leaf or clump of moss that shelters it from weather. The clutch of two eggs is incubated by the female, who first breeds in her second year. The incubation period and time to fledging are not known.[7]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to amethyst-throated sunangel on xeno-canto
Vocalization

The amethyst-throated sunangel's call is "a short, buzzy, dry trill...repeated at intervals."[7]
Status

The IUCN has assessed the amethyst-throated sunangel as being of Least Concern. Its population size is not known but is believed to be stable.[1] It is fairly common overall, locally abundant, and occurs in several protected areas.[7]
References

BirdLife International (2016). "Amethyst-throated Sunangel Heliangelus amethysticollis". IUCN Red List of Threatened Species. 2016: e.T60859997A95164192. doi:10.2305/IUCN.UK.2016-3.RLTS.T60859997A95164192.en. Retrieved 20 January 2022.
"Appendices | CITES". cites.org. Retrieved 2022-01-14.
Gill, F.; Donsker, D.; Rasmussen, P. (July 2021). "IOC World Bird List (v 12.1)". doi:10.14344/IOC.ML.11.2. Retrieved January 15, 2022.
HBW and BirdLife International (2020) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip [.xls zipped 1 MB] retrieved May 27, 2021
Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022
del Hoyo, J., I. Heynen, N. Collar, P. F. D. Boesman, G. M. Kirwan, and C. J. Sharpe (2020). Amethyst-throated Sunangel (Heliangelus amethysticollis), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.amtsun1.01 retrieved January 20, 2022