Haliaeetus leucoryphus

Haliaeetus leucoryphus (*)

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Accipitriformes

Familia: Accipitridae
Subfamilia: Haliaeetinae
Genus: Haliaeetus
Species: Haliaeetus leucoryphus
Name

Haliaeetus leucoryphus (Pallas, 1771)
Synonyms

Aquila leucorypha (protonym)

References

Reise durch verschiedene Provinzen des Russischen Reichs 1: 454.

Vernacular names
বাংলা: কুড়া
brezhoneg: Morerer Pallas
čeština: Orel páskovaný
Deutsch: Bindenseeadler
English: Pallas's Fish Eagle
Esperanto: Strivosta maraglo
español: Pigargo de Pallas
suomi: Aromerikotka
français: Pygargue de Pallas
magyar: Szalagos rétisas
日本語: キガシラウミワシ
norsk: Båndhavørn
ଓଡ଼ିଆ: ବଇରିଗଞ୍ଜନ
polski: Bielik wschodni
русский: Орлан-долгохвост
ไทย: อินทรีทะเลหัวนวล
中文: 玉带海雕

Pallas's fish eagle (Haliaeetus leucoryphus), also known as Pallas's sea eagle or band-tailed fish eagle, is a large, brownish sea eagle. It breeds in the east Palearctic in Kazakhstan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Mongolia, China, India, Nepal, Bangladesh, Myanmar and Bhutan. It is listed as Endangered on the IUCN Red List.[1] It is partially migratory, with Central Asian birds wintering among the southern Asian birds in northern India, and also further west to the Persian Gulf.[2]
Description
Pallas's fish eagle in flight at Jim Corbett National Park

The Pallas's fish eagle has a light sandy-brown hood and a whitish face. The wings are darker brown and the back rufous. The long, slender wings (particularly slender for a sea eagle) are rather dark brown underneath. The tail is black with a wide, distinctive white stripe. Juveniles are overall darker, cooler brown with no band on the tail but with several pale areas on the wing, including the underwing coverts and inner primaries. This results in underwings that have a white band in young fish eagles. It takes until the 4th year or so to obtain adult plumage. Among related species,, it mainly overlaps in range with the quite dissimilar much shorter winged and slightly smaller grey-headed fish eagle and scarcely with the larger, bulkier and much broader winged white-tailed eagle, which also seldom resembles the coloring of the Pallas's.[3][4] This fairly large species measures 72 to 85 cm (28 to 33 in) in length with a wingspan of 180–215 cm (5 ft 11 in – 7 ft 1 in).[2] Females are generally reported to weigh 2.1 to 3.7 kg (4.6 to 8.2 lb), with this sample of nine averaging 3.2 kg (7.1 lb), and are slightly larger than males at 2.03 to 3.3 kg (4.5 to 7.3 lb), in ten that weighed an average of 2.6 kg (5.7 lb).[4][5][6] However, in some cases Pallas's fish eagles have been reported to weigh as much as 4 to 5.5 kg (8.8 to 12.1 lb) and span as much as much as 240 cm (7 ft 10 in).[7][8] Thus, their size falls just slightly under the large northern sea eagles (i.e. bald, white-tailed and Steller's) and broadly similar to slightly larger than the sea eagles of more tropical central distribution.[4]
Behaviour and ecology
Diet

Its diet consists primarily of large freshwater fish. However, Pallas's fish eagle seems to have a broadly opportunistic diet much like better known sea eagles.[4] They also regularly prey upon birds, especially water birds. Other prey reportedly can include mammals, including leporids and rodents, frogs, reptiles, including snakes and terrapins, insects (i.e. wasp larvae) and carrion.[4][9] Their prey in Bangladesh were reported to consist of mrigal carp, rohu, catla, ilish, Chitala chitala, Mystus cavasius, Ompok bimaculatus and Wallago attu.[9] Known water bird prey can include pheasant-tailed jacana, lesser whistling duck, ferruginous duck, common pochard and little grebe.[9] Pallas's fish eagles are known to attack quite large (i.e. about their own size or somewhat larger) adult water birds including greylag geese, bar-headed geese and Demoiselle crane by assaulting them on the surface of the water and then flying off with the kill. Since the greylag goose species is slightly heavier than the eagle, this is one of the greatest weight-lifting feats ever recorded for a flying bird.[9] Another case of lifting a great load was recorded at the Yamuna River in north-central India, where an eagle captured a huge carp and flew with the struggling fish very low over the water, before dropping it in response to gunfire. The carp was found to have weighed 6.5 kg (14 lb), which is probably roughly twice the weight of the eagle carrying it.[10] How they interact with other raptors is not well understood. However, observations of wintering Pallas's fish eagles in Bharatpur suggest they may dominate several other eagle species, select the highest perches and have the highest daily food intake, including over similarly sized eagles such as eastern imperial eagles and steppe eagles.[11]
Taxonomy
Pallas's fish eagle eggs

Aquila leucorypha was the scientific name proposed by Peter Simon Pallas in 1771, who first described a Pallas's fish eagle that he encountered during his travels in eastern Siberia.[12] In the 19th century, it was also placed in the genera Falco and Haliaeetus by different authors.[13]

This species is the hardest-to-place sea-eagle. Among the species of its genus, it has no close living relatives. mtDNA cytochrome b sequence data is unable to reliably suggest a phylogenetic place for it among the sea-eagles. However, some information can be drawn from the molecular data, and especially from morphology and biogeography:

This species retains the ancestral dark eye, bill, and talons of the first sea-eagles, shared with the older tropical lineage. It is peculiar insofar as it has a black band at the end of the tail in adult birds, similar to juvenile Madagascar fish eagles (which look like a smaller, darker version of this bird, but are not very closely related). Its distribution indicates that this species evolved fairly independently of other sea-eagle lineages, but the molecular data tentatively suggests it is possibly closer to the Holarctic species. It diverged from its common ancestor with other species soon after the Holarctic and the tropical lineages split. Dependent on the interpretation of a possible Early Oligocene sea-eagle fossil from Egypt, this happened either at the very start or the end of the Oligocene, somewhere between 34 and 25 mya.[14]
Distribution and habitat
Juvenile Pallas's fish eagle in Bangladesh

The Pallas's fish eagle occurs in Russia, Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan, Mongolia, China, India, Nepal, Bangladesh, Myanmar and Bhutan.[1]
Conservation

The Pallas's fish eagle is listed as Endangered on the IUCN Red List. The global population is estimated at less than 2,500 individuals. Besides direct persecution, humans contribute to the decline of this species through habitat degradation, pollution, and draining or overfishing lakes. In India, the spread of water hyacinth in lakes possibly makes finding prey difficult for the Pallas's fish eagle.[1] Its large range is deceptive, as Pallas's fish eagle is rare and isolated throughout its territory and may not breed in large areas of it.[15]
References

BirdLife International (2018). "Haliaeetus leucoryphus". IUCN Red List of Threatened Species. 2018: e.T22695130A131934599. doi:10.2305/IUCN.UK.2018-2.RLTS.T22695130A131934599.en. Retrieved 19 November 2021.
del Hoyo, Elliott & Sargatal 1994.
Robson, C. (2000). A Field Guide to the Birds of South-east Asia. New Holland, London, UK.
Ferguson-Lees, J., Christie, D. A. (2001). Raptors of the World. Boston, New York: Houghton Mifflin Harcourt. ISBN 0-618-12762-3.
Brazil, M. (2019). Field Guide to the Birds of East Asia. Bloomsbury Publishing.
Dunning Jr, J. B. (2007). CRC Handbook of Avian Body Masses, 2nd Edition. CRC Press.
Steele, M. (2017). Where in the World are Pallas's Fish Eagles? Migration and Ecology of Haliaeetus leucoryphus in Asia. University of Arkansas.
Paz, U. (1987). The birds of Israel. Stephen Greene Press.
Orta, J., D. A. Christie, G. M. Kirwan, and C.J. Sharpe (2020). Pallas's Fish-Eagle (Haliaeetus leucoryphus), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA.
Wood, G. (1983). The Guinness Book of Animal Facts and Feats. ISBN 978-0-85112-235-9.
Prakash, V. (1988). The general ecology of raptors in Keoladeo National Park, Bharatpur (Doctoral dissertation, Ph. D. thesis. Bombay University, Mumbai, India).
Pallas, P. S. (1771). "Aquila leucorypha". Reise durch verschiedene Provinzen des russischen Reichs in einem ausführlichen Auszuge. Volume 2. Frankfurt und Leipzig: J. G. Fleischer. pp. 454–455.
Bree, C. R. & Fawcett, B. (1859). "Pallas's sea eagle". A history of the birds of Europe, not observed in the British Isles. Vol. 1. London: Groombridge and Sons. pp. 75–79.
Wink, M.; Heidrich, P.; Fentzloff, C. (1996). "A mtDNA phylogeny of sea eagles (genus Haliaeetus) based on nucleotide sequences of the cytochrome b gene" (PDF). Biochemical Systematics and Ecology. 24 (7–8): 783–791. doi:10.1016/S0305-1978(96)00049-X.
del Hoyo, J.; Elliott, A.; Sargatal, J., eds. (1994). Handbook of the Birds of the World. Vol. 2. Barcelona: Lynx Ediciones. ISBN 84-87334-15-6.