Colaptes rivolii

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Piciformes

Familia: Picidae
Subfamilia: Picinae
Genus: Colaptes
Species: Colaptes rivolii
Subspecies: C. r. atriceps - C. r. brevirostris - C. r. meridae - C. r. quindiuna - C. r. rivolii - C. r. zuliensis
Name

Colaptes rivolii (Boissonneau, 1840)
Synonyms

Picus rivolii (protonym)
Piculus rivolii (Boissonneau, 1840)
Colaptes elegans G.R. Gray
Hypoxanthus rivolii (Boissonneau, 1840)

References

Boissonneau, A. 1840. Nouvelle espèce du genre Picus. Revue Zoologique par la Société Cuviérienne 3: 36–37. BHL Reference page. p. 36 BHL

Vernacular names
English: Crimson-mantled Woodpecker
español: Carpintero candela; carpintero carmesí
français: Pic de Rivoli

The crimson-mantled woodpecker (Colaptes rivolii) is a species of bird in subfamily Picinae of the woodpecker family Picidae. It is found in Bolivia, Colombia, Ecuador, Peru, and Venezuela.[3]
Taxonomy and systematics

The crimson-mantled woodpecker was originally described as Picus Rivolii.[4] It was later placed in genus Hypoxanthus that was still later merged into Piculus; since about 2007 it has been moved into Colaptes by taxonomic systems.[5][3][6][7][8]

The American Ornithological Society, the International Ornithological Committee, and the Clements taxonomy recognize these six subspecies of crimson-mantled woodpecker:[8][3][7]

C. r. quindiuna (Chapman, 1923)
C. r. zuliensis (Aveledo & Peréz, 1989)
C. r. rivolii (Boissonneau, 1840)
C. r. meridae (Chapman, 1923)
C. r. brevirostris (Taczanowski, 1874)
C. r. atriceps (Sclater and Salvin, 1876)

BirdLife International's Handbook of the Birds of the World (HBW) treats C. r. atriceps as a separate species, the black-crowned woodpecker.[6]

The specific epithet rivolii honors French ornithologist François Victor Masséna, third Duke of Rivoli and third Prince of Essling.[9]

This article follows the six-subspecies model.
Description

The crimson-mantled woodpecker is 23 to 26 cm (9.1 to 10 in) long and weighs 85 to 112 g (3.0 to 4.0 oz). Males and females have the same plumage except on their heads. Adult males of the nominate subspecies C. r. rivolii have a red crown. Their face is mostly pale yellow with a dark red malar stripe and a black chin and throat. Adult females have a black crown and malar stripe. Both sexes have a crimson red nape and mantle; their rump and uppertail coverts are black. Their wings are bronzy red and their tail is black. Their underparts are yellow with black and red scallops on the breast and sometimes black spots on the belly. Their bill is black, their iris red-brown to brown, and the legs pale gray. Juveniles are generally duller than adults and have no red on the breast; the male's crown is blacker.[10]

The other subspecies of crimson-mantled woodpecker vary little from the nominate with the exception of C. r. atriceps. Males of that subspecies have a black crown that is more extensive than the female's. Both sexes have no red on the breast and their wings are less red than those of the nominate.[10]
Distribution and habitat

The subspecies of crimson-mantled woodpecker are found thus:[3][10]

C. r. quindiuna, Colombia's Cordillera Central
C. r. zuliensis, the Serranía del Perijá on the Colombia-Venezuela border
C. r. rivolii, Colombia's Cordillera Oriental into northwestern Venezuela
C. r. meridae, the Andes of northwestern Venezuela between Trujillo and Táchira
C. r. brevirostris, the Andes from southwestern Colombia south through Ecuador into central Peru
C. r. atriceps, the Andes of southeastern Peru and western and central Bolivia (The range map omits this subspecies)

The crimson-mantled woodpecker inhabits the interior and edges of humid montane forest and the lower edge of the páramo zone.[10] In Ecuador it mostly ranges at elevations between 2,000 and 3,300 m (6,600 and 10,800 ft).[11]
Behavior
Feeding

The crimson-mantled woodpecker forages at any level of the forest, typically on moss- and lichen-covered limbs but also on trunks. Its primary food is small arthropods, especially ants; it also feeds on fruit. It usually forages singly or in pairs, but does regularly join mixed species feeding flocks.[10] It shuns the introduced shrub gorse.[12]
Breeding

The crimson-mantled woodpecker's breeding season has not been defined but appears to vary geographically. One author surmised that it spans June to November in Peru and another suggested that (possibly species-wide) it includes February to March. It nests in tree cavities but nothing else is known about its breeding biology.[10]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to crimson-mantled woodpecker on xeno-canto
Vocal and non-vocal sounds

What is thought to be the song of the crimson-mantled woodpecker's most widespread subspecies brevirostris is "a series of fairly monotone, rapid metallic notes...kee'r'r-ker'r-ke'r'r. The apparent song of atriceps is a "descending low churr...grr'r'r'r'l". The species also makes "a rising REE?" call. The crimson-mantled woodpecker drums, but not frequently.[10]
Status

The IUCN follows HBW taxonomy and so has assessed the crimson-mantled sensu stricto and black-crowned woodpeckers separately. Both are rated as being of Least Concern. The population size of neither is known but both are believed to be stable. No immediate threats to either have been identified.[1][2] "Human activity has little direct effect on the Crimson-mantled Woodpecker, other than the local effects of habitat destruction."[10]
References

BirdLife International (2016). "Crimson-mantled Woodpecker Colaptes rivolii". IUCN Red List of Threatened Species. 2016: e.T22726390A94920901. doi:10.2305/IUCN.UK.2016-3.RLTS.T22726390A94920901.en. Retrieved 28 January 2023.
BirdLife International (2016). "Black-crowned Woodpecker Colaptes atriceps". IUCN Red List of Threatened Species. 2016: e.T22726398A94921093. doi:10.2305/IUCN.UK.2016-3.RLTS.T22726398A94921093.en. Retrieved 28 January 2023.
Gill, F.; Donsker, D.; Rasmussen, P., eds. (August 2022). "Woodpeckers". IOC World Bird List. v 12.2. Retrieved January 9, 2023.
Boissonneau, Auguste (February 1840). Guérin-Mémeville, F.-E. (ed.). "Nouvelle espèce du genre Pic". Travaux inédits. Revue Zoologique par la Société Cuvierienne (in French): 36–37. Retrieved January 28, 2023.
Benz, Brett W.; Robbins, Mark B.; Peterson, A. Townsend (2006). "Evolutionary history of woodpeckers and allies (Aves: Picidae): Placing key taxa on the phylogenetic tree". Molecular Phylogenetics and Evolution. 40 (2): 389–399. Bibcode:2006MolPE..40..389B. doi:10.1016/j.ympev.2006.02.021. PMID 16635580.
HBW and BirdLife International (2022) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 7. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v7_Dec22.zip retrieved December 13, 2022
Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022
Jobling, James A. (2010). Helm Dictionary of Scientific Bird Names. London, UK: Christopher Helm. pp. 336–337. ISBN 978-1-4081-2501-4.
Schulenberg, T. S. (2020). Crimson-mantled Woodpecker (Colaptes rivolii), version 1.0. In Birds of the World (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.crmwoo2.01 retrieved January 28, 2023
Ridgely, Robert S.; Greenfield, Paul J. (2001). The Birds of Ecuador: Field Guide. Vol. II. Ithaca: Cornell University Press. p. 334. ISBN 978-0-8014-8721-7.
Amaya-Villarreal, Ángela (November 2010). "Effects of Gorse (Ulex europaeus) on the birds of a high Andean forest edge". Ornitología Colombiana. 10: 11–25.