Tetraphalerus

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Archostemata
Superfamilia: Cupedoidea

Familia: Ommatidae
Subfamilia: Ommatinae
Tribus: Ommatini
Genus: Tetraphalerus
Species (28): T. bruchi – T. wagneri – †T. antiquus – †T. bontsaganensis – †T. brevicapitis – †T. brevis – †T. collaris – †T. curtinervis – †T. decorosus – †T. fentaiensis – †T. incertus – †T. glabratus – †T. grandis – †T. laetus – †T. largicoxa – †T. latus – †T. lentus – †T. longicollis – †T. macilentus – †T. maximus – †T. mongolicus – †T. notatus – †T. okhotensis – †T. penalveri – †T. ponomarenkoi – †T. surrectus – †T. tenuipes – †T. verrucosus
Name

Tetraphalerus Waterhouse, 1901
Type species: Tetraphalerus wagneri Waterhouse, 1901
Fixation: monotypy

Synonyms

?Longaevicupes Ren, 1995

NOTE: name missing from ION

Monticupes Ren, 1995

NOTE: name missing from ION

Odontomma Tan, Ren & Ge, 2006
Authority for synonymy: Kirejtshuk et al., 2010

References
Primary references

Waterhouse, C.O. 1901: Two new genera of Coleoptera belonging to the Cupesidae and Prionidae. Annals and magazine of natural history (7), 7(42): 520–523. DOI: 10.1080/00222930108678509 BHL

Additional references

Atkins, M.D. 1963: The Cupedidae of the World. Canadian entomologist, 95(2): 140–162. [Publication date: 'February 1963', sourced from first page of article] DOI: 10.4039/Ent95140-2 [160]
Beutel, R.G.; Ge, S.-Q.; Hörnschemeyer, T. 2008: On the head morphology of Tetraphalerus, the phylogeny of Archostemata and the basal branching events in Coleoptera. Cladistics, 24: 270–298. DOI: 10.1111/j.1096-0031.2007.00186.x
Kirejtshuk, A.G., Ponomarenko, A.G., Prokin, A.A., Chang, H., Nikolajev, G.V. & Ren, D. 2010. Current knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (northeast China). Acta Geologica Sinica - English Edition 84(4): 783–792. DOI: 10.1111/j.1755-6724.2010.00253.x Paywall. Reference page.
Ponomarenko, A.G.; Martinez-Delclos, X. 2000: New beetles (Insecta: Coleoptera) from the Lower Cretaceous of Spain. Acta Geologica Hispanica, 35: 47–52.
Ren, D. 1995: Fossil insects. In Faunae and stratigraphy of the Jurassic-Cretaceous in Beijing and adjacent areas, Beijing. Seismic, Beijing.
Soriano, C.; Delclos, X. 2006: New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family. Acta Palaeontologica Polonica, 51(1): 185–200. PDF
Tan, J.; Ren, D.; Shih, C. 2007: New beetles (Insecta: Coleoptera: Archostemata) from the Late Mesozoic of North China. Annales zoologici, 57: 231–247. PDF

Links

ION search for Tetraphalerus
Nomenclator Zoologicus search for Tetraphalerus
Tree of Life Web Project 2006. Tetraphalerus. Version 10 February 2006 (temporary) in The Tree of Life Web Project

Tetraphalerus is a genus of beetles in the family Ommatidae, It is currently known from two extant species native to South America and several fossil species from the Jurassic and Cretaceous of Asia.
Systematics

[1]

According to Kirejtshuk, 2020.[2]

†Tetraphalerus brevis Ponomarenko, 1964 Karabastau Formation, Kazakhstan, Oxfordian
Tetraphalerus bruchi Heller, 1913, recent, Northern Argentina
†Tetraphalerus collaris Ponomarenko, 1997 Dzun-Bain Formation, Mongolia, Aptian
†Tetraphalerus glabratus Ponomarenko, 1997 Togo-Khuduk Formation, Mongolia, Bajocian/Bathonian
†Tetraphalerus largicoxa Lin Qibin, 1986 Guanyintan Formation, China Toarcian/Aalenian
†Tetraphalerus mongolicus Ponomarenko, 1986 Gurvan-Eren Formation, Mongolia, Aptian
†Tetraphalerus notatus Ponomarenko, 1997 Dzun-Bain Formation, Mongolia, Aptian
Tetraphalerus wagneri Waterhouse, 1901, recent, South America

Formerly assigned species

†Tetraphalerus antiquus Ponomarenko, 1964 assigned to Allophalerus
†Tetraphalerus bontsaganensis Ponomarenko, 1997 assigned to Allophalerus
†Tetraphalerus brevicapitis Pononarenko & Martinez-Delclos, 2000 considered indeterminate by Kirejtshuk, 2020
†Tetraphalerus fentaiensis (Ren, 1995) assigned to Monticupes
†Tetraphalerus grandis Ponomarenko, 1964 assigned to Omma
†Tetraphalerus incertus Ponomarenko, 1969 assigned to Allophalerus
†Tetraphalerus laetus Lin Qibin, 1976 assigned to Zygadenia
†Tetraphalerus latus Tan et al. 2007 assigned to Allophalerus
†Tetraphalerus lindae Jarzembowski et al. 2017 assigned to Bukhkalius
†Tetraphalerus longicollis Ponomarenko, 1997 assigned to Omma
†Tetraphalerus maximus Ponomarenko, 1968 assigned to Allophalerus
†Tetraphalerus okhotensis Ponomarenko in Cromov et al., 1993 assigned to Allophalerus
†Tetraphalerus oligocenicus (Crowson, 1962) assigned to Tetraphalerites
†Tetraphalerus tenuipes Ponomarenko, 1964 assigned to Allophalerus
†Tetraphalerus verrucosus Ponomarenko, 1966 assigned to Allophalerus
†Tetraphalerus surrectus (Ren, 1995) assigned to Monticupes

References

Ommatidae Species List Archived 2012-10-23 at the Wayback Machine at Joel Hallan’s Biology Catalog. Texas A&M University. Retrieved on 11 May 2012.
Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. doi:10.3390/geosciences10020073. ISSN 2076-3263.