Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Elateriformia
Superfamilia: Elateroidea
Familia: Rhagophthalmidae
Genera: ?Cydistus – Dioptoma – Diplocladon – Dodecatoma – ?Falsophrixothrix – Menghuoius – Mimoochotyra – Pseudothilmanus – Rhagophthalmus
Name
Rhagophthalmidae
References
selected references
Kawashima, I.; Lawrence, J.F.; Branham, M.A. 2010: 4.14. Rhagophthalmidae Olivier, 1907. Pp. 135-140 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755 DOI: 10.1515/9783110911213.135
Kundrata, R.; Bocak, L. 2011: Redescription and relationships of Pseudothilmanus Pic (Coleoptera: Rhagophthalmidae)—a long-term neglected glow-worm beetle genus from the Himalayas. Zootaxa, 2794: 57–62. Preview
Li, X.-Y.; Xie, M.; Liang, X.-C. 2008: Rhagophthalmid - a special group of fireflies. Chinese bulletin of entomology, 45(3): 494–498. [In Chinese, with English abstract] full article (PDF)
Wittmer, W.; Ohba, N. 1994: Neue Rhagophthalmidae (Coleoptera) aus China und benachbarten Landern. Japanese journal of entomology, 62(2): 341–355. CiNii
Links
Rhagophthalmidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
The Rhagophthalmidae are a family of beetles within the superfamily Elateroidea. Members of this beetle family have bioluminescent organs on the larvae, and sometimes adults, and are closely related to the Phengodidae (glowworm beetles), though historically they have been often treated as a subfamily of Lampyridae, or as related to that family.[1] Some recent evidence suggested that they were the sister group to the Phengodidae, and somewhat distantly related to Lampyridae, whose sister taxon was Cantharidae,[2] but more reliable genome-based phylogenetics placed (Rhagophthalmidae + Phengodidae) as the sister group to the Lampyridae.[3][4]
Whatever their relationships may be, Rhagophthalmidae are distributed in the Old World, and little is known of their biology. Females are usually wingless and look like larvae, but have an adult beetle's eyes, antennae and legs; in the genus Diplocladon, they resemble larvae even more, with small light organs on all trunk segments. Larvae and females live in soil and litter and are predaceous; males may be attracted to lights at night.[5][6]
Genera
Bicladodrilus
Bicladon
Dioptoma
Diplocladon
Dodecatoma
Falsophrixothrix
Mimoochotyra
Mondodrilus
Pseudothilmanus[7]
Reductodrilus
Rhagophthalmus
References
Stanger-Hall KF, Lloyd JE, Hillis DM (October 2007). "Phylogeny of North American fireflies (Coleoptera: Lampyridae): implications for the evolution of light signals". Molecular Phylogenetics and Evolution. 45 (1): 33–49. doi:10.1016/j.ympev.2007.05.013. PMID 17644427.
Bocak L, Motyka M, Bocek M, Bocakova M (2018-03-14). Chiang TY (ed.). "Incomplete sclerotization and phylogeny: The phylogenetic classification of Plastocerus (Coleoptera: Elateroidea)". PLOS ONE. 13 (3): e0194026. Bibcode:2018PLoSO..1394026B. doi:10.1371/journal.pone.0194026. PMC 5851614. PMID 29538419.
Kusy D, Motyka M, Bocek M, Vogler AP, Bocak L (November 2018). "Genome sequences identify three families of Coleoptera as morphologically derived click beetles (Elateridae)". Scientific Reports. 8 (1): 17084. Bibcode:2018NatSR...817084K. doi:10.1038/s41598-018-35328-0. PMC 6244081. PMID 30459416.
Branham MA, Wenzel JW (2001). "The evolution of bioluminescence in cantharoids (Coleoptera: Elateroidea)". Florida Entomologist. 84 (4): 565–586. doi:10.2307/3496389. ISSN 1938-5102. JSTOR 3496389.
Branham MA, Wenzel JW (February 2003). "The origin of photic behavior and the evolution of sexual communication in fireflies (Coleoptera: Lampyridae)". Cladistics. 19 (1): 1–22. doi:10.1111/j.1096-0031.2003.tb00404.x. PMID 34905865. S2CID 46266960.
Lawrence JF, Hastings AM, Dallwitz MJ, Paine TA, Zurcher EJ (October 2005). Elateriformia (Coleoptera): descriptions, illustrations, identification, and information retrieval for families and subfamilies: Lampyridae (Report). version of 9. Archived from the original on 27 November 2010.
Kundrata R, Bocak L (March 2011). "Redescription and relationships of Pseudothilmanus Pic (Coleoptera: Rhagophthalmidae)—a long-term neglected glow-worm beetle genus from the Himalayas". Zootaxa. 2794 (1): 57–62. doi:10.11646/ZOOTAXA.2794.1.4.
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