Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Supercohort: Polyneoptera
Cohort: Dictyoptera
Ordo: Blattodea
Superfamilia: Corydioidea
Familia: Nocticolidae
Genera (9): Alluaudellina – Cardacopsis – Cardacus – Helmablatta – Metanocticola – Nocticola – Pholeosilpha – Spelaeoblatta – Typhloblatta – Typhloblattodes
[source: Blattodea Species File (Version 5.0/5.0)]
In synonymy (2): Alluaudella – Cardax
Name
Nocticolidae Bolívar, 1892
References
Bolivar I., 1892. Voyage de M E Simon aux iles Philippines (mars et avril 1890). Études sur les Arthropodes cavernicoles de l'île de Luzon. Orthoptères. Annales de la Société Entomologique de France.
Beccaloni, G. & Eggleton, P. 2013. Order Blattodea. Zootaxa 3703(1): 46–48. DOI: 10.11646/zootaxa.3703.1.10 Open access. Reference page. [see p. 47, biodiversity: 9 genera, 32 species]
Vidlička, Ľ., Vršanský, P., Kúdelová, T., Kúdela, M., Deharveng, L. & Hain, M. 2017. New genus and species of cavernicolous cockroach (Blattaria, Nocticolidae) from Vietnam. Zootaxa 4232(3): 361–375. DOI: 10.11646/zootaxa.4232.3.5. Reference page.
Nocticolidae is a small family in the order Blattodea (cockroaches). It consists of only 32 known species in 9 genera. They are found in Africa, Asia and Australia. Most live in cave habitats, although a few are associated with termites. Cave adapted species are known from the Cenomanian aged Burmese amber, making them the oldest extant cavernicolous organisms.[1]
Genera
The family includes the following genera:
Alluaudellina Chopard, 1932
Cardacopsis Karny, 1924
Cardacus Strand, 1928
Metanocticola Roth, 1999
Nocticola Bolívar, 1892
Pholeosilpha Chopard, 1958
Spelaeoblatta Bolívar, 1897
Typhloblatta Chopard, 1924
Typhloblattodes Chopard, 1946
Crenocticola[2] Li and Huang, 2019 Burmese amber, Cenomanian
Mulleriblattina[1] Sendi et al, 2020 Burmese amber, Cenomanian
References
Sendi, Hemen; Vršanský, Peter; Podstrelená, Lenka; Hinkelman, Jan; Kúdelová, Tatiana; Kúdela, Matúš; Vidlička, Ľubomír; Ren, Xiaoyin; Quicke, Donald L.J. (February 2020). "Nocticolid cockroaches are the only known dinosaur age cave survivors". Gondwana Research. 82: 288–298. doi:10.1016/j.gr.2020.01.002.
Li, Xin-Ran; Huang, Diying (February 2020). "A new mid-Cretaceous cockroach of stem Nocticolidae and reestimating the age of Corydioidea (Dictyoptera: Blattodea)". Cretaceous Research. 106: 104202. doi:10.1016/j.cretres.2019.104202. ISSN 0195-6671.
Blattodea Species File: Nocticolidae
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