Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Elateriformia
Superfamilia: Elateroidea
Familia: Phengodidae
Subfamilia: Mastinocerinae
Genera: Akamboja – Brasilocerus – Cenophengus – ?Cephalophrixothorax – Decamastinocerus – Distremocephalus – Eurymastinocerus – Euryopa – Euryognathus – Howdenia – Mastinocerus – Mastinomorphus – Mastinowittmerus – Neophengus – Nephromma – Oxymastinocerus – Paraphrixothrix – Paraptorthodius – Phrixothrix – Pseudomastinocerus – Ptorthodiellus – Ptorthodius – Spangleriella – ?Steneuropa – Stenophrixothrix – Taximastinocerus
References
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M. & Silveira, L.F.L. 2017. Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae). Zootaxa 4306(4): 501–523. DOI: 10.11646/zootaxa.4306.4.3. Paywall. Reference page.
Vega-Badillo, V., Morrone, J.J. & Zaragoza-Caballero, S. 2021. Revision of the genus Cenophengus LeConte, 1881 (Coleoptera, Phengodidae), with the description of four new species, new geographic records and a new synonymy. Zookeys 1068ː 73–148. DOI: 10.3897/zookeys.1068.70295 Open access Reference page.
Zaragoza-Caballero, S. 2010: Paraphrixothrix ecuadoranus, a new genus and species of Phengodidae (Coleoptera: Mastinocerinae) from Ecuador. Pan-Pacific entomologist, 86(4): 105–110. DOI: 10.3956/2007-40.1
The Mastinocerinae are a subfamily of phengodid beetles (Phengodidae). It contains mostly Central and South American species.[1] They are also known as "railroad worms".
Genera[1]
Akamboja Roza, Quintino, Mermudes & Silveira, 2017, 5 spp.
Brasilocerus Wittmer, 1963, 9 spp.
Cenophengus LeConte, 1881, 23 spp.
Cephalophrixothrix Wittmer, 1976, 3 spp.
Decamastinocerus Wittmer, 1988, 2 spp.
Distremocephalus Wittmer, 1976, 11 spp.
Eurymastinocerus Wittmer, 1976, 8 spp.
Euryognathus Wittmer, 1976, 2 spp.
Euryopa Gorham, 1881, 8 spp.
Howdenia Howdenia Wittmer, 1976, 10 spp.
Mastinocerus Solier, 1849
subgenus Mastinocerus Solier, 1849, 18 spp.
subgenus Paramastinocerus Wittmer, 1976, 8 spp.
Mastinomorphus Wittmer, 1976, 15 spp.
Mastinowittmerus Zaragoza-Caballero, 1984, 2 spp.
Neophengus Wittmer, 1976, 3 spp.
Nephromma Wittmer, 1976, 2 spp.
Oxymastinocerus Wittmer, 1963, 9 spp.
Paraphrixothrix Zaragoza-Caballero, 2010, 1 sp.
Paraptorthodius Schaeffer, 1904, 3 spp.
Phrixothrix Olivier, 1909, 17 spp.
Pseudomastinocerus Wittmer, 1963, 9 spp.
Ptorthodiellus Wittmer, 1976, 2 spp.
Ptorthodius Gorham, 1881, 3 spp.
Spangleriella Wittmer, 1988, 1 sp.
Steneuryopa Wittmer, 1986, 1 sp.
Stenophrixothrix Wittmer, 1963, 19 spp.
Taximastinocerus Wittmer, 1963, 18 spp.
References
Kundrata, Robin; Blank, Stephan M; Prosvirov, Alexander S; Sormova, Eliska; Gimmel, Matthew L; Vondráček, Dominik; Kramp, Katja (2019-10-19). "One less mystery in Coleoptera systematics: the position of Cydistinae (Elateriformia incertae sedis) resolved by multigene phylogenetic analysis". Zoological Journal of the Linnean Society. Oxford University Press (OUP). doi:10.1093/zoolinnean/zlz104. ISSN 0024-4082.
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