Leptotes

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Lycaenidae
Subfamilia: Polyommatinae
Tribus: Polyommatini
Genus: Leptotes
Species (alphabetic): L. adamsoni – L. andicola – L. babaulti – L. bathyllos – L. brevidentatus – L. callanga – L. casca – L. cassioides – L. cassius – L. delalande – L. durrelli – L. jeanneli – L. lamasi – L. lybas – L. mandersi – L. marginalis – L. marina – L. mayottensis – L. parrhasioides – L. perkinsae – L. pirithous – L. plinius – L. pulchra – L. pyrczi – L. rabefaner – L. sanctithomae – L. socotranus – L. trigemmatus – L. webbianus

Species (Afrotropical):

L. adamsoni – L. babaulti – L. brevidentatus – L. casca – L. cassioides – L. durrelli – L. jeanneli – L. mandersi – L. marginalis – L. mayottensis – L. pirithous – L. pulchra – L. pyrczi – L. rabefaner – L. sanctithomae – L. socotranus – L. webbianus

Species (Oriental):

L. plinius

Species (Australasian):

L. lybas

Species (Neotropical):

L. andicola – L. bathyllos – L. callanga – L. cassius – L. delalande – L. lamasi – L. marina – L. parrhasioides – L. perkinsae – L. trigemmatus
Name

Leptotes Scudder, 1876: 124.

Type species: Lycaena theonus Lucas, 1857.
Synonyms

Cyclyrius Butler, [1897]: 830.
Type species: Polyommatus webbianus Brullé, 1840, by original designation.
Syntarucus Butler, [1901]: 929.
Type species: Papilio telicanus Lang, 1789, by original designation.
Syntarucoides Kaye, 1904: 190.
Type species: Papilio cassius Cramer, [1775], by original designation.
Langia Tutt, 1906: 314 (praeocc. Moore, 1872).
Type species: Papilio telicanus Lang, 1789, by original designation.
Raywardia Tutt, 1908: 414, replacement name for Langia Tutt, 1906.

References
Primary references

Scudder, S.H. 1875–76. Synonymic list of the butterflies of North America, north of Mexico. Bulletin of the Buffalo Society of Natural Sciences 2: 233–269; 3: 98–129; Pt.1 Nymphales, Pt.2 Rurales. Reference page.
Butler, A.G. [1897]. On two collections of Lepidoptera made by Mr. Richard Crawshay in Nyasa-Land. Proc. zool. Soc. Lond. 1896(4): 817–850, pls 41,42. BHL, plates. Reference page.
Butler, A.G. [1901]. On two consignments of butterflies collected by Mr. Richard Crawshay in the Kikuyu country of British East Africa in 1899 and 1900. Proceedings of the Zoological Society of London 1900: 911–946, Pl.58. text, Plate. Reference page.
Kaye, W.J. 1904. A catalogue of the Lepidoptera Rhopalocera of Trinidad. Trans. ent. Soc. Lond. 52(2): 159–228, pls 17,18. text, plates. Reference page.
Tutt, J.W. 1905–1906. A Natural History of the British Lepidoptera Vol 8, Butterflies (1): 479pp, 20 pls. London, Swan Sonnenschein & Co. BHL. Reference page.
Tutt, J.W. 1907–1908. A Natural History of the British Lepidoptera Vol 9, Butterflies (2): 495pp, 28 pls. London, Swan Sonnenschein & Co. BHL Reference page.

Additional references

Bálint, Z. & Johnson, K. 1995. Species diagnostics of the genus Leptotes in continental South America (Lepidoptera, Lycaenidae). Reports of the Museum of Natural History, University of Wisconsin (Stevens Point) 44: 1–24. PDF. Reference page.
Bridges, C.A. 1994. Catalogue of the Family-Group, Genus:-Group and Species-Group Names of the Riodinidae & Lycaenidae (Lepidoptera) of the World. Bridges, Urbana, Illinois. BHL Reference page.
Fric, Z.F., Maresova, J., Kadlec, T., Tropek, R., Pyrcz, T.W. & Wiemers, M. 2019. World travellers: phylogeny and biogeography of the butterfly genus Leptotes (Lepidoptera: Lycaenidae). Systematic Entomology 44(3): 652–665. DOI: 10.1111/syen.12349 Reference page.
Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol. 5A. Association for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439p. Reference page.
Libert, M. 2011. Note sur le genre Leptotes Scudder, 1876 (Lepidoptera, Lycaenidae). Lambillionea 111(1): 53–66. Reference page.
Libert, M., Baliteau, L. & Baliteau, S. 2011. Deux nouveaux Lycaenidae du Cap-Vert (Lepidoptera). Bulletin de la Société entomologique de France 116(1): 63–67. PDF. Reference page.
Schwartz, A. & Johnson, K. 1992. Two new butterflies (Lepidoptera: Lycaenidae) from Cuba. Caribbean Journal of Science 28(3-4): 149–157. PDF. Reference page.
Stempffer, H. 1967. The genera of the African Lycaenidae (Lepidoptera: Rhopalocera). Bulletin of the British Museum (Natural History), entomology. Supplement 10: 322pp, 1pl. BHL Reference page.
Williams, M.C. 2007. Checklist of Afrotropical Papilionoidea and Hesperioidea. 7th Edn. (April 2007). Downloaded from Afrotropical Butterflies. Reference page.

Vernacular names
English: Zebra Blues
中文: 細灰蝶屬

Leptotes is a butterfly genus in the family Lycaenidae. They are commonly known as zebra blues in reference to their zebra-striped undersides.

The genus Cyclyrius was recently synonymized with Leptotes and its two species were thus moved to this genus.[1]

Species

The genus can be divided in several distinct geographic groups which often correspond to clades:[1]

Afrotropical and Palaearctic species:

Leptotes adamsoni Collins & Larsen, 1991 – Adamson's zebra blue
Leptotes babaulti (Stempffer, 1935) – Babault's zebra blue
Leptotes brevidentatus (Tite, 1958) – Tite's zebra blue
Leptotes casca (Tite, 1958)
Leptotes cassioides (Capronnier, 1889)
Leptotes durrelli Fric, Pyrcz & Wiemers, 2019
Leptotes jeanneli (Stempffer, 1935)
Leptotes mandersi (Druce, 1907)
Leptotes marginalis (Stempffer, 1944) – black-bordered zebra blue
Leptotes mayottensis (Tite, 1958)
Leptotes pirithous (Linnaeus, 1767) – Lang's short-tailed blue, common zebra blue
Leptotes pulcher (Murray, 1874) – beautiful zebra blue
Leptotes pyrczi Libert, 2011
Leptotes rabefaner (Mabille, 1877)
Leptotes sanctithomae (Sharpe, 1893)
Leptotes socotranus (Ogilvie-Grant, 1899)
Leptotes webbianus (Brullé, 1839) – Canary blue

Indomalayan species:

Leptotes plinius (Fabricius, 1793) – zebra blue, plumbago blue

Australasian species:

Leptotes lybas (Godart, [1824])

Neotropical species:

Leptotes andicola (Godman & Salvin, 1891)
Leptotes bathyllos Tessmann, 1928
Leptotes callanga (Dyar, 1913)
Leptotes cassius (Cramer, [1775]) – Cassius blue, tropical striped blue
Leptotes delalande Bálint & Johnson, 1995
Leptotes lamasi Bálint & Johnson, 1995
Leptotes marina (Reakirt, 1868) – marine blue, striped blue
Leptotes parrhasioides (Wallengren, 1860) – Galapagos blue
Leptotes perkinsae Kaye, 1931
Leptotes trigemmatus (Butler, 1881)

Leptotes pirithous
Footnotes

Fric et al., 2019.

References

Brower, Andrew V.Z. (2008): Tree of Life Web Project – Leptotes Scudder 1876. The Zebra Blues. Version of 2008-MAY-19. Retrieved 2009-NOV-29.
Savela, Markku (2009): Markku Savela's Lepidoptera and some other life forms – Leptotes. Version of 2009-MAY-24. Retrieved 2009-NOV-29.
Fric, Z.F., Maresova, J., Kadlec, T., Tropek, R., Pyrcz, T.W. & Wiemers, M., 2019. World travellers: phylogeny and biogeography of the butterfly genus Leptotes (Lepidoptera: Lycaenidae). Systematic Entomology 44(3): 652–665. doi:10.1111/syen.12349