Elcanidae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Orthoptera
Subordo: Ensifera
Infraordo: Elcanidea
Superfamilia: †Elcanoidea
Familia: Elcanidae
Subfamiliae: Archelcaninae – †Baisselcaninae - Elcaninae

Unassigned Elcanidae: †Archelcana - †Burmelcana - †Cratoelcana - †Elcanonympha –

†Hispanelcana - †Parelcana - †Probaisselcana - †Sibelcana - Sinoelcana – †Synelcana
Name

Elcanidae Handlirsch, 1906
References

Handlirsch, A. 1906–1908. Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch für Paläontologen und Zoologen. Leipzig: W. Engelmann. 1430 pp. [1906: pp. 1–640, 1907: pp. 641–1120, 1908: pp. 1120–1430]. BHL Reference page.
Gu, J-J., Tian, H., Wang, J-Y., Zhang, W-Z., Ren, D. & Yue, Y-L. 2020. A world key to the genera of Elcanidae (Insecta, Orthoptera), with a Jurassic new genus and species of Archelcaninae from China. ZooKeys, 954: 65–74. DOI: 10.3897/zookeys.954.52088 Open access Reference page.
Heads, S.W., Thomas, M.J. & Wang, Y-N. 2018. A new genus and species of Elcanidae (Insecta: Orthoptera) from Cretaceous Burmese amber. Zootaxa 4527(4): 575–580. DOI: 10.11646/zootaxa.4527.4.8 Paywall Reference page.
Peñalver, E.; Grimaldi, D.A. 2010: Latest occurrences of the Mesozoic family Elcanidae (Insecta: Orthoptera), in Cretaceous amber from Myanmar and Spain. Annales de la Société entomologique de France (n.s.), 46: 88–99. ISSN: 0037-9271 [not seen]
Tian, H., Gu, J-J., Yin, X-C. & Ren, D. 2019. The first Elcanidae (Orthoptera, Elcanoidea) from the Daohugou fossil bed of northeastern China. ZooKeys, 897: 19–28. DOI: 10.3897/zookeys.897.37608 Open access Reference page.

Taxonomy

†subfamily Archelcaninae Gorochov, 2011[6]
†Archelcana Sharov, 1968[7]
†Archelcana durnovaria Whalley, 1985[8] – Charmouth Mudstone Formation, England, Early Jurassic (Sinemurian)
†Archelcana ornata Zherikhin, 1985[9] – Cheremkhovskaya Formation, Russia, Early Jurassic (Toarcian)
†Archelcana shurabica Sharov, 1968 – Sagul Formation, Kyrgyzstan, Toarcian
Indeterminate remains attributed to the genus are also known from the latest Triassic of Germany[10]
†Jeholelcana Fang et al. 2018b[11]
†Jeholelcana yanensis Fang et al. 2018b – Yixian Formation, China, Early Cretaceous (Aptian)
†Parelcana Handlirsch, 1906[12]
†Parelcana anglicana Handlirsch, 1939[13] – Blue Lias, United Kingdom, Early Jurassic (Hettangian)
†Parelcana dubia Handlirsch, 1939 – United Kingdom, Late Triassic (Rhaetian)
†Parelcana pulchmacula Tian et al., 2019[14] – Daohugou, China, Middle Jurassic (Callovian)
†Parelcana tenuis Handlirsch, 1906 – Green Series, Germany, Toarcian
†Sibelcana Gorochov, 1990[15]
†Sibelcana rossica Gorochov, 1990 – Glushkovo Formation, Russia, Late Jurassic (Tithonian)
†Sibelcana transbaicalica Gorochov, 1990 – Ukurei Formation, Russia, Tithonian
†Synelcana Zessin, 1988[16]
†Synelcana muelleri Zessin, 1988 – Green Series, Germany, Toarcian
†subfamily Elcaninae Handlirsch, 1906
†Cratoelcana Martins-Neto, 1991[17] – Crato Formation, Brazil, Aptian
†Cratoelcana damianii Martins-Neto, 1991
†Cratoelcana zessini Martins-Neto, 1991
†Ellca Kočárek, 2020[1]
†Ellca nevelka Kočárek, 2020 – Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
†Eubaisselcana Gorochov, 1986[18] – Gurvan-Eren Formation, Mongolia, Aptian
†Eubaisselcana mongolica Gorochov, 1986
†Eubaisselcana sharovi Gorochov, 1986
†Minelcana Gorochov et al., 2006[19]
†Minelcana dubia (Giebel 1856)[20] – Lulworth Formation, United Kingdom, Early Cretaceous (Berriasian)
†Minelcana membranacea Gorochov et al., 2006 – Durlston Formation, United Kingdom, Berriasian
†Panorpidium Westwood, 1854[21]
†Panorpidium bimaculatum Gorochov et al., 2006[19] – Weald Clay, United Kingdom, Early Cretaceous (Barremian)
†Panorpidium maculosum Zhou et al. 2022[22] Burmese amber, Myanmar, Cenomanian
†Panorpidium proximum Gorochov et al., 2006 – Durlston Formation, United Kingdom, Berriasian
†Panorpidium parvum Gorochov et al., 2006 – Weald Clay, United Kingdom, Early Cretaceous (Hauterivian)
†Panorpidium sibirica Sharov, 1968[7] – Zaza Formation, Russia, Aptian
†Panorpidium spica Kim et al., 2021[3] – Jinju Formation, South Korea, Early Cretaceous (Albian)
†Panorpidium tessellatum Westwood, 1854 – Durlston Formation, United Kingdom, Berriasian
†Panorpidium yixianensis Fang et al., 2015[23] – Yixian Formation, China, Aptian
†Probaisselcana Gorochov, 1989[24]
†Probaisselcana cretacea Gorochov et al., 2006[19] – Weald Clay, United Kingdom, Hauterivian
†Probaisselcana euryptera Tian et al., 2019[2] – Yixian Formation, China, Aptian
†Probaisselcana karatavica Sharov, 1968[7] – Karabastau Formation, Kazakhstan, Late Jurassic (Oxfordian)
Incertae sedis
†Burmelcana Peñalver & Grimaldi, 2010[25]
†Burmelcana longirostris Peñalver & Grimaldi, 2010 – Burmese amber, Myanmar, Cenomanian
†Cascadelcana Fang et al., 2018a[4]
†Cascadelcana virginiana Fang et al., 2018a – Cow Branch Formation, North Carolina, Late Triassic (Norian)
†Cenoelcanus Schubnel et al., 2020[26]
†Cenoelcanus menatensis Schubnel et al., 2020 – Menat Formation, France, Paleocene
†Elcanonympha Heads et al., 2018[27]
†Elcanonympha diana Heads et al., 2018 – Burmese amber, Myanmar, Cenomanian
†Hispanelcana Peñalver & Grimaldi, 2010 – Álava amber, Escucha Formation, Spain, Albian
†Hispanelcana alavensis Peñalver & Grimaldi, 2010
†Hispanelcana arilloi Peñalver & Grimaldi, 2010
†Hispanelcana lopezvallei Peñalver & Grimaldi, 2010
†Longioculus Poinar et al., 2007[28]
†Longioculus burmensis Poinar et al., 2007 – Burmese amber, Myanmar, Cenomanian
†Hukawnelca Uchida, 2021[29]
†Hukawnelca gracile Uchida, 2021 – Burmese amber, Myanmar, Cenomanian
†Caelielca Uchida, 2021[29]
†Caelielca spinocrus Uchida, 2021 – Burmese amber, Myanmar, Cenomanian
†Monitelcana Xu et al., 2022[5]
†Monitelcana penalveri Xu et al., 2022 – Burmese amber, Myanmar, Cenomanian

References

Kočárek P (December 2020). "A diminutive elcanid from mid-Cretaceous Burmese amber, Ellca nevelka gen. et sp. nov., and the function of metatibial spurs in Elcanidae (Orthoptera)". Cretaceous Research. 116: 104574. doi:10.1016/j.cretres.2020.104574.
Tian H, Gu JJ, Huang F, Zhang H, Ren D (July 2019). "A new species of Elcaninae (Orthoptera, Elcanidae) from the Lower Cretaceous Yixian Formation at Liutiaogou, Inner Mongolia, NE China, and its morphological implications". Cretaceous Research. 99: 275–280. doi:10.1016/j.cretres.2019.03.010.
Kim, Do-Yoon; Lee, Mirinae; Nam, Gi-Soo; Park, Tae-Yoon S. (April 2021). "The first orthopteran fossils from the Lower Cretaceous (Albian) Jinju Formation of Korea: ethological implications for elcanids". Cretaceous Research. 125: 104843. doi:10.1016/j.cretres.2021.104843. ISSN 0195-6671.
Fang Y, Muscente AD, Heads SW, Wang B, Xiao S (2018-05-25). "The earliest Elcanidae (Insecta, Orthoptera) from the Upper Triassic of North America". Journal of Paleontology. 92 (6): 1028–1034. doi:10.1017/jpa.2018.20. ISSN 0022-3360. S2CID 133821611.
Xu, Chunpeng; Luo, Cihang; Jarzembowski, Edmund A.; Fang, Yan; Wang, Bo (2022-03-28). "Aposematic coloration from Mid-Cretaceous Kachin amber". Philosophical Transactions of the Royal Society B: Biological Sciences. 377 (1847): 20210039. doi:10.1098/rstb.2021.0039.
Gorochov AV (2011). "New family-group taxa in the infraclass Polyneoptera (Insecta): making the names, proposed early, available" (PDF). Zoosystematica Rossica. 20: 159–160. doi:10.31610/zsr/2011.20.1.159.
Sharov AG (1968). "Filogeniya ortopteroidnykh nasekomykh" [Phylogeny of orthopteroid insects]. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR (in Russian). 118: 1–216.
Whalley PE (1985). "The systematics and palaeogeography of the Lower Jurassic insects of Dorset, England". Bulletin of the British Museum (Natural History) Geology. 39 (3): 107–189.
Zherikhin VV (1985). "Yurskie pryamokrylye yuzhnoy Sibiri i zapadnoy Mongolii (Gryllida=Orthoptera)" [Jurassic Orthoptera of southern Siberia and western Mongolia (Grillida = Orthnoptera).]. Yurskie Nasekomye Sibiri I Mongolii (in Russian): 171–184.
Barth G, Ansorge J, Brauckmann C (December 2011). "First record of the genus Ipsvicia (Hemiptera: Ipsviciidae) outside Gondwana - an Australian genus from the Upper Triassic of Germany". Polish Journal of Entomology / Polskie Pismo Entomologiczne. 80 (4): 645–657. doi:10.2478/v10200-011-0050-z. ISSN 0032-3780. S2CID 84850244.
Fang Y, Heads SW, Wang H, Zhang H, Wang B (August 2018). "The first Archelcaninae (Orthoptera, Elcanidae) from the Cretaceous Jehol Biota of Liaoning, China". Cretaceous Research. 86: 129–134. doi:10.1016/j.cretres.2018.02.008.
Handlirsch A (1906). "Die Fossilen Insekten und die Phylogenie der Rezenten Formen, parts I-IV." [The fossil insects and the phylogeny of the recent forms]. Ein Handbuch fur Palaontologen und Zoologen [A manual for palaontologists and zoologists] (in German). pp. 1–640.
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Tian H, Gu JJ, Yin XC, Ren D (2019-12-09). "The first Elcanidae (Orthoptera, Elcanoidea) from the Daohugou fossil bed of northeastern China". ZooKeys (897): 19–28. doi:10.3897/zookeys.897.37608. PMC 6914707. PMID 31857785.
Gorochov V (1990). "Orthopterans, Gryllida, in Pozdne-Mezozoyskie Nasekomye Vostochnogo Zabaykal'ya" [Orthnopterans, Grillida, in Late Mesozoic Insects of Eastern Transbaikalia]. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta [USSR Academy of Sciences, Proceedings of the Paleontological Institute] (in Russian). 239: 210–214.
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Gorochov AV, Jarzembowski EA, Coram RA (2006-10-01). "Grasshoppers and crickets (Insecta: Orthoptera) from the Lower Cretaceous of southern England". Cretaceous Research. 27 (5): 641–662. doi:10.1016/j.cretres.2006.03.007. ISSN 0195-6671.
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Zhou, Qian; Xu, Chunpeng; Jarzembowski, Edmund A.; Xiao, Chuantao (2022-08-01). "A new species of Elcanidae (Insecta: Orthoptera) from mid-Cretaceous Kachin amber". Cretaceous Research. 136: 105226. doi:10.1016/j.cretres.2022.105226. ISSN 0195-6671.
Fang Y, Wang B, Zhang H, Wang H, Jarzembowski EA, Zheng D, Zhang Q, Li S, Liu Q (January 2015). "New Cretaceous Elcanidae from China and Myanmar (Insecta, Orthoptera)". Cretaceous Research. 52: 323–328. doi:10.1016/j.cretres.2014.05.004.
Gorochov AB (1989). "Novye taksony prjamokrylyx semejstv Bintoniellidae, Xenopteridae, Permelcanidae, Elcanidae, i Vitimiidae (Orthoptera, Ensifera) iz mezozoja azii" [New taxa of the Orthopteran families Bintoniellidae, Xenopteridae, Permelcanidae, Elcanidae, and Vitimiidae (Orthoptera, Ensifera) from the Mesozoic Asia]. Vestnik Zoologii (in Russian). 1989 (4): 20–27.
Peñalver E, Grimaldi DA (2010-01-01). "Latest occurrences of the Mesozoic family Elcanidae (Insecta: Orthoptera), in Cretaceous amber from Myanmar and Spain". Annales de la Société Entomologique de France. N.S. 46 (1–2): 88–99. doi:10.1080/00379271.2010.10697641. ISSN 0037-9271. S2CID 84115545.
Schubnel, Thomas; Desutter, Laure; Garrouste, Romain; hervé, sophie; Nel, André (2020). "Paleocene of Menat Formation, France, reveals an extraordinary diversity of orthopterans and the last known survivor of a Mesozoic Elcanidae". Acta Palaeontologica Polonica. 65. doi:10.4202/app.00676.2019. ISSN 0567-7920.
Heads SW, Thomas MJ, Wang Y (December 2018). "A new genus and species of Elcanidae (Insecta: Orthoptera) from Cretaceous Burmese amber". Zootaxa. 4527 (4): 575–580. doi:10.11646/zootaxa.4527.4.8. PMID 30651491.
Poinar G, Gorochov AV, Buckley R (2007). "Longioculus burmensis, n. gen., n. sp. (Orthoptera: Elcanidae) in Burmese amber". Proceedings of the Entomological Society of Washington. 109: 649–655.
Uchida, Kei (2021-11-17). "Two new genera of the Elcanidae (order: Orthoptera) from middle Cretaceous Burmese amber of northern Myanmar". Cretaceous Research: 105092. doi:10.1016/j.cretres.2021.105092. ISSN 0195-6671.