Dorcus

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Scarabaeoidea

Familia: Lucanidae
Subfamilia: Lucaninae
Genus: Dorcus
Species: D. alexisi – D. amamianus – D. antaeus – D. arfakianus – D. arrowi – D. bandaensis – D. bashanus – D. bolanus – D. branaungi – D. brevis – D. capricornis – D. carinulatus – D. cervulus – D. chayuensis – D. cuonaensis – D. curvidens – D. cylindricus – D. daedalion – D. davidis – D. derelictus – D. detanii – D. donckieri – D. egregius – D. fujii – D. fuliginosus – D. furusui – D. fuscescens – D. ghiliani – D. gongshanus – D. grandis – D. haitschunus – D. hamidi – D. hansi – D. hansteini – D. hirticornis – D. hopei – D. hyperion – D. immundus – D. insolitus – D. intermedius – D. intricatus – D. japonicus – D. kamijoi – D. katsurai – D. kawamurai – D. kesiniae – D. kikunoae – D. kirchneri – D. koreanus – D. kyawi – D. lachnosternus – D. lacroixi – D. laevidorsis – D. lhoba – D. lineatopunctatus – D. linwenhsini – D. linzhiensis – D. liyingbingi – D. lvbu – D. macleayi – D. maryi – D. mattisi – D. meeki – D. menba – D. mencius – D. mineti – D. mirabilis – D. miwai – D. mizunumai – D. monpa – D. montivagus – D. multicavus – D. musimon – D. myinti – D. nepalensis – D. niasicus – D. niedorfi – D. nitidus – D. nosei – D. parallelipipedus – D. parallelus – D. pemakoi – D. peyronis – †D. primigenius – D. prochazkai – D. rama – D. ratiocinativus – D. rectus – D. reichei – D. ritsemae – D. rubrofemoratus – D. rugosus – D. saiga – D. schenklingi – D. semenowi – D. sewertzowi – D. sinensis – D. striatipennis – D. submolaris – D. sukkiti – D. suturalis – D. taiwanicus – D. tanakai – D. taoi – D. tatsuhikoi – D. tenuihirsutus – D. ternatensis – D. tianlongi – D. tityus – D. townesi – D. tsunodai – D. ursulus – D. velutinus – D. vicinus – D. vidam – D. wardi – D. wickhami – D. wui – D. yaksha – D. yamadai – D. yuukae – D. zhangmuensis
Name

Dorcus MacLeay, 1819: 111

Type species: Scarabaeus parallelipipedus Linnaeus, 1758.

Synonymy

Ditomoderus Parry, 1864
Type species: Ditomoderus mirabilis Parry, 1864
Durelius Houlbert, 1915
Type species: Dorcus derelicticus Parry, 1862
Dynodorcus Didier, 1931
Type species: Dorcus antaeus Hope, 1842 (Hope 1842)
Eurydorcus Didier, 1931
Type species: Dorcus reichei Hope, 1842 (Hope 1842)
Eurytrachellelus Didier, 1931
Type species: Dorcus tityus Hope, 1842 (Hope 1842)
Eurytrachelus Thomson, 1862
Type species: Dorcus tityus Hope, 1842 (Hope 1842)
Goniodorcus Didier, 1931
Type species: Eurytrachelus coranus Gestro, 1881
Hemisodorcus Thomson, 1862
Type species: Lucanus nepalensis (Hope, 1831)
Hemisodorcus (Paradorcus) Maes, 1990
Macrodorcas Motschulsky, 1861
Macrodorcus Felsche, 1898
Nipponodorcus Nomura & Kurosawa, 1960
Telodorcus Didier, 1931
Velutinodorcus Maes, 1992

References
Primary references

Macleay, W.S. 1819. Horae entomologicae: or essays on the annulose animals. Containing geneal observations on the geography, manners, and natural affinities of the Insects which compose the genus Scarabaeus of Linnaeus; to which are added a few incidental remarks on the genera Lucanus and Hister of the same author. With an appendix and plates. Vol. 1. Part 1. London: S. Bagster, xxx + [2] + 160 pp., 3 pls. BHL Reference page. [original description: p. 111]

Additional references

Araya, K. 2012: Notes on Eurytrachelus dorcoides Nagel, 1939 from Sumatra, a junior synonym of Dorcus rama (Boileau, 1897). Kogane, (13): 147–150. [not seen] Reference page.
Araya, K., Hosoya, T. 2005. Molecular phylogeny of the Dorcus velutinus species-group (Coleoptera: Lucanidae) inferred from the mitochondrial 16S rRNA gene sequences with the special reference to the taxonomic status of its Japanese and Taiwanese members. Elytra, Tokyo 33(2): 523–530.
Baba, M. 2005. A new species of the genus Dorcus Macleay, 1819 from Northeast Burma. Gekkan-Mushi (414): 44–48.
Baba, M. 2008. A new species of the genus Dorcus MacLeay, 1819 from Obi Is., Indonesia. Gekkan-Mushi (450): 38–40.
Baba, M. 2008. A new species of the genus Dorcus MacLeay, 1819 from northwest India. Gekkan-Mushi (453): 42–44.
Bartolozzi, L. & Werner, K. 2004. Illustrated catalogue of the Lucanidae from Africa and Madagascar. Taita Publishers: Czech Republic. ISBN 80-902734-7-5
Fujita, H. 2009. Two new species and a new subspecies of Lucanid-beetles from Southern Vietnam. Gekkan-Mushi (462): 2–5. (Japanese)
Hosoguchi 2004. A new species of the genus Dorcus (Coleoptera, Lucanidae) from northern India. Gekkan-Mushi (402): 30–31.
Hosoguchi 2006. A new species of the genus Dorcus (Coleoptera, Lucanidae) from Enggano Is. Indonesia. Gekkan-Mushi (426): 20–22.
Ikeda, H. 2000. A new species of the genus Dorcus (Coleoptera, Lucanidae) from northern Vietnam. Gekkan-Mushi 350: 28–29. Reference page.
Jang, Y.-C. & Kawai, S. 2008. A new species of the Dorcus velutinus-group (Coleoptera, Lucanidae) from Korea. Kogane (9): 103–106.
Kim, S.I. & Kim, J-I. 2010. Review of family Lucanidae (Insecta: Coleoptera) in Korea with the description of one new species. Entomological Research 40: 55–81. DOI: 10.1111/j.1748-5967.2009.00263.x
Maeda, K. 2009. Three new species of the genera Lucanus, Rhaetulus and Dorcus (Coleoptera, Lucanidae) from central Vietnam. Gekkan-Mushi (457): 35–40. (Japanese)
Muret, P. & Drumont, A. 1999. Description d'une nouvelle espece de Dorcus Macleay, endemique de Chypre (Coleoptera, Scarabaeoidea, Lucanidae) [Description of a new species of Dorcus MacLeay, endemic to Cyprus (Coleoptera, Scarabaeoidea, Lucanidae).] Lambillionea 99: 484–488.
Nagai, S. 2002. Notes on some SE Asian stag-beetles (Coleoptera, Lucanidae), with descriptions of several new taxa. Gekkan-Mushi (372): 11–14.
Nagai, S. & Maeda, K. 2009. Four new species of stag-beetles (Coleoptera, Lucanidae) from Myanmar. Gekkan-Mushi (462): 46–49. (Japanese)
Okuda, N. & Maeda, T. 2014. Two new species and a new subspecies of the family Lucanidae (Coleoptera) from Arunachal Pradesh, northeastern India. Gekkan-Mushi (522): 13–17.
Paulsen, M.J. 2010: Stag beetles of the genus Dorcus MacLeay in North America (Coleoptera, Lucanidae). In: Ratcliffe, B.& Krell, F.-T. (eds) Current advances in Scarabaeoidea research. ZooKeys 34: 199–207. DOI: 10.3897/zookeys.34.292
Schenk, K.-D. 2008. Contribution to the knowledge of the stag beetles of Asia (Coleoptera, Lucanidae) and description of several new taxa. Beetles World (1) [in German with English abstract] [Publication date: 'March, 2008', sourced from first page of article]
Schenk, K.D. 2016. Description of Nigidius grosseri from Uganda and Dorcus maryi from New Guinea (Coleoptera, Lucanidae). Beetles World 13: 1-5. Reference page.
Wang, C-B. & Zhou, C. 2019. Dorcus tianlongi, a new species from central China (Coleoptera: Lucanidae: Lucaninae). Zootaxa 4691(5): 575–588. DOI: 10.11646/zootaxa.4691.5.9 Paywall Reference page.

Dorcus is a genus of beetles in the stag beetle family Lucanidae. Of the over 100 species, most occur in Asia and India; two are found in southern Europe, and two species are from North America. Previously, specimens with serriform teeth on the mandibles and sable pigment were called Serrognathus whereas specimens with but a singular or multiple bulky notches on the mandibles and lustrous sable pigmentation were called Dorcus.[1]
Species

The following species are recognised in the genus Dorcus:[2]

Dorcus alcides
Dorcus alexisi Muret & Drumont, 1999 - Cyprus
Dorcus amamianus (Nomura, 1964)
Dorcus antaeus Hope, 1842
Dorcus arfakianus (Lansberge, 1880) - Papua New Guinea
Dorcus arrowi (Boileau, 1911) - southern and south-eastern Asia
Dorcus bandaensis Okuda, 2000 - Indonesia
Dorcus bashanus Huang & Chen, 2013 - China
Dorcus bolanus Nagel, 1928 - Papua New Guinea
Dorcus branaungi Nagai, 2000 - Myanmar, China
Dorcus brevis (Say, 1825) - North America
Dorcus bucephalus
Dorcus capricornis (Möllenkamp, 1909) - Solomon Islands
Dorcus carinulatus Nagel, 1941 - temperate Asia
Dorcus cervulus (Boileau, 1901) - Vietnam, Thailand, China
Dorcus chayuensis Huang & Chen, 2017 - China
Dorcus consentaneus
Dorcus cuonaensis Huang & Chen, 2013 - China
Dorcus curvidens (Hope, 1840)
Dorcus cylindricus Thomson, 1862 - India
Dorcus daedalion (Didier & Séguy, 1953) - China, south-eastern Asia
Dorcus davidis (Fairmaire, 1887) - China, Mongolia
Dorcus derelictus Parry, 1862 - India
Dorcus detanii Mizunuma, 1994 - Indonesia
Dorcus donckieri (Boileau, 1898)
Dorcus egregius (Möllenkamp, 1898) - New Guinea, Solomon Islands
Dorcus fujii Nagai & Maeda, 2010 - Vietnam
Dorcus fuliginosus (Didier, 1928) - India
Dorcus furusui Baba, 2008 - Indonesia
Dorcus fuscescens Didier, 1931 - Indonesia
Dorcus ghiliani (Gestro, 1881) - Indonesia
Dorcus gongshanus Huang & Chen, 2013 - China
Dorcus grandis Didier, 1926
Dorcus haitschunus (Didier & Séguy, 1952) - China, North Korea, South Korea
Dorcus hamidi Ikeda, 2005 - Indonesia
Dorcus hansi Schenk, 2008 - China
Dorcus hansteini (Albers, 1889)
Dorcus hirticornis (Jakovlev, 1896)
Dorcus hopei (Saunders, 1854)
Dorcus hyperion (Boileau, 1899)
Dorcus immundus Arrow, 1938 - India
Dorcus insolitus (Didier, 1931) - Indonesia
Dorcus intermedius (Gestro, 1881) - Oceanea
Dorcus intricatus (Lacroix, 1981) - China
Dorcus japonicus Nakane & Makino, 1985 - Japan
Dorcus kamijoi Fujita, 2009 - Vietnam
Dorcus katsurai Ikeda, 2000 - Vietnam, China
Dorcus kawamurai Fujita, 2010 - India
Dorcus kesiniae (Choeyjanta, Pathomwattananurak & Chanthapoon, 2018) - eastern Asia
Dorcus kikunoae Hosoguchi, 2004 - India
Dorcus kirchneri Schenk, 2008 - Indonesia
Dorcus koreanus Jang & Kawai, 2008
Dorcus kyanrauensis (Miwa, 1934)
Dorcus kyawi Nagai & Maeda, 2009 - Myanmar, China
Dorcus lachnosternus (de Lisle, 1972)
Dorcus lacroixi (Bomans, 1973) - Malaysia
Dorcus laevidorsis Fairmaire, 1888
Dorcus lhoba Huang & Chen, 2013 - China
Dorcus lineatopunctatus (Hope, 1831)
Dorcus linwenhsini Huang & Chen, 2013 - China
Dorcus linzhiensis Huang, Chen, Tao & Xiao, 2020 - China
Dorcus liyingbingi Huang & Chen, 2013 - China
Dorcus lvbu Huang & Chen, 2013 - China
Dorcus macleayi (Hope, 1845) - India
Dorcus maryi Schenk, 2016 - Papua New Guinea
Dorcus mattisi Schenk, 2022
Dorcus meeki Boileau, 1906
Dorcus menba Huang & Chen, 2013 - China
Dorcus mencius (Kriesche, 1935) - China
Dorcus mineti de Lisle, 1974
Dorcus mirabilis (Parry, 1864) - Malaysia
Dorcus miwai Benesh, 1936 - Taiwan and temperate Asia
Dorcus mizunumai Fujita, 2010 - Indonesia
Dorcus mochizukii (Miwa, 1937)
Dorcus monpa Okuda & Maeda, 2014 - India
Dorcus montivagus (Lewis, 1883)
Dorcus multicavus Mizunuma, 1994
Dorcus musimon Gené, 1836 - southern Europe and North Africa
Dorcus myinti Nagai & Maeda, 2009 - Myanmar, China
Dorcus nepalensis (Hope, 1831) - India
Dorcus niasicus (Nonfried, 1895) - Indonesia
Dorcus niedorfi Schenk, 2013 - Myanmar
Dorcus nitidus Kirsch, 1877 - Indonesia, Papua New Guinea
Dorcus nosei Nagai, 2000 - Myanmar, China
Dorcus parallelipipedus (Linnaeus, 1758) - lesser stag beetle; Palearctic, Mexico (introduced)
Dorcus parallelus (Say, 1824) - North America
Dorcus pemakoi Huang, Okuda, Maeda & Chen, 2017 - China
Dorcus peyronis Reiche & Saulcy, 1856 - Europe, Middle East
†Dorcus primigenius Deichmüller, 1881 - Eocene of Czech Republic
Dorcus prochazkai Schenk, 2003 - Iran
Dorcus rama (Boileau, 1897) - Indonesia
Dorcus ratiocinativus Westwood, 1871 - India
Dorcus rectus (Motschulsky, 1857) - temperate Asia
Dorcus reichei Hope, 1842 - Indomalaya, China
Dorcus ritsemae Oberthür & Houlbert, 1914
Dorcus rubrofemoratus (Snellen van Vollenhoven, 1865)
Dorcus rugosus Boileau, 1904 - India
Dorcus saiga (Olivier, 1789)
Dorcus schenklingi (Möllenkamp, 1913) - Taiwan and temperate Asia
Dorcus seguyi
Dorcus semenowi (Jakovlev, 1900) - China
Dorcus sewertzowi Semenov, 1891 - western and southwestern Asia
Dorcus sinensis (Boileau, 1899)
Dorcus striatipennis (Motschulsky, 1861) - Japan, Russia Far East
Dorcus submolaris (Hope, 1845) - Indomalaya
Dorcus sukkiti Fukinuki, 2004 - Myanmar, China
Dorcus suturalis Westwood, 1871 - Afghanistan, Pakistan, India, Thailand
Dorcus taiwanicus Nakane & Makino, 1985 - Taiwan and temperate Asia
Dorcus tanakai Nagai, 2002 - China, Vietnam
Dorcus taoi Huang & Chen, 2020 - China
Dorcus tatsuhikoi Fujii & Okuda, 2020 - Vietnam
Dorcus tenuihirsutus Kim & Kim, 2010 - South Korea, China
Dorcus ternatensis Thomson, 1862 - Indonesia
Dorcus tianlongi Wang & Zhou, 2019 - China
Dorcus tityus Hope, 1842 - Bangladesh, India
Dorcus townesi (de Lisle, 1972) - Philippines
Dorcus tsunodai Fujita, 2009 - Vietnam
Dorcus ursulus Arrow, 1938 - India, Bhutan, Myanmar, Thailand, China
Dorcus velutinus Thomson, 1862 - China, Indomalaya
Dorcus vicinus Saunders, 1854 - China
Dorcus vidam (Nguyen & Schenk, 2015) - Vietnam
Dorcus wardi Arrow, 1943 - China, Myanmar
Dorcus wickhami (Waterhouse, 1894) - Australia
Dorcus wui Huang & Chen, 2013 - China
Dorcus yaksha Gravely, 1915
Dorcus yamadai (Miwa, 1937) - Taiwan and temperate Asia
Dorcus yuukae Bartolozzi & Nagai, 2017 - Philippines
Dorcus zhangmuensis Huang & Chen, 2013 - China
BOLD:ADJ7517 (Dorcus sp.)

References

Brett C. Ratcliffe (2002). "Lucanidae". In Ross H. Arnett Jr. & Michael C. Thomas (ed.). American Beetles. CRC Press. p. 8.
"Dorcus Macleay, 1819". www.gbif.org. Retrieved 2024-05-20.