Cryptochetidae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Brachycera
Infraordo: Muscomorpha
Sectio: Schizophora
Subsectio: Acalyptrata
Superfamilia: Carnoidea

Familia: Cryptochetidae
Genera (2 + 1†): Cryptochetum – Librella – †Phanerochaetum

[source: Catalogue of Life: 2013 Annual Checklist, except fossils]
Name

Cryptochetidae Brues & Melander, 1932

References
Links

Tree of Life Web Project. 2007. Cryptochetidae. Version 29 November 2007 (temporary). http://tolweb.org/Cryptochetidae/10588/2007.11.29 in The Tree of Life Web Project, http://tolweb.org/

Vernacular names
中文: 隐毛蝇科

The Cryptochetidae are a small family of tiny flies (generally 2 to 4 mm long). Some twenty to thirty species are known. Generally they are metallic blue black, stoutly built, with the head broad and high and with clear wings.[1] Like other species in the superfamily Lonchaeoidea, the Cryptochetidae have antennae with a cleft in the second segment. Unlike practically all Schizophora however, they lack an arista, or if they do have one, it is too small to distinguish with any confidence. The family name refers to this unusual distinction; "Cryptochetidae" literally means "those with hidden bristles". The adult flies also are unusual among insects in that they have only a single pair of abdominal spiracles — this is not a serious physiological challenge in such small insects.

Again in resemblance to other Lonchaeoidea, the Cryptochetidae do not have more than one proclinate orbital bristle on each side. The frons is densely setulose. The costa has a break at the end of the subcosta. The sixth abdominal sternite in the male is symmetrical and it has an 8th tergite. In the female the seventh tergite and sternite are fused, and the eighth segment is elongated.

The larvae are of biological and economic interest, being endoparasitoids of coccids. In Cryptochetum iceryae, which parasitizes Icerya, there are four larval instars. The first instar is sac-like and lacks both trophi and tracheae but at the caudal end it bears a pair of finger-like processes. The caudal end of the digestive tract is closed. During subsequent instars the caudal processes grow longer and become filamentous; in the final instar they are much longer than the whole body. These filaments probably are respiratory organs. Only a few species of Cryptochetidae have been described, and most of those occur in tropical countries. At one time they were allocated to the Agromyzidae but now are regarded as a separate family.[2]
Genera

These three genera belong to the family Cryptochetidae:

Cryptochetum Rondani, 1875[3]
Librella McAlpine, 1976[4]
Phanerochaetum Hennig, 1965[5]

References

McAlpine, J. F. et al (eds) Manual of Nearctic Diptera. Monographs No. 27 & No. 28 Biosystematics Research Institute, Ottawa. Ontario Research Branch. Pub: Agriculture Canada Vol 1 1981 & Vol 2 1987
Richards, O. W.; Davies, R.G. (1977). Imms' General Textbook of Entomology: Volume 1: Structure, Physiology and Development Volume 2: Classification and Biology. Berlin: Springer. ISBN 0-412-61390-5.
Rondani, Camillo (1875). "Species Italicae ordinis Dipterorum (Muscaria Rndn.) collectae et observatae. Stirps XXIII. Agromyzinae". Bullettino della Società Entomologica Italiana. 7: 166–191. Retrieved 7 November 2020.
McAlpine, D.K. (1976). "A new genus of flies possibly referable to Cryptochetidae (Diptera, Schizophora)". Austral. Ent. Mag. 3 (3): 45–56.
Hennig, Willi (1965). "Die Acalyptratae des Baltischen Bernsteins und ihre Bedeutung fur die Erforschung der phylogenetischen Entwicklung dieser Dipteren-Gruppe". Stuttgarter Beiträge zur Naturkunde. 145: 215. Retrieved 7 November 2020.