Coccidulini

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Coccinelloidea

Familia: Coccinellidae
Subfamilia: Coccinellinae
Tribus: Coccidulini
Genera (85): Acoccidula – Adoxellus – Ambrocharis – Anisorhizobius – Apolinus – Apseudoscymnus – Auladoria – Aulis – Axinoscymnus – Azya – Botynella – Bucolus – Carinoscymnus – Chnoodes – Clitostethus – Coccidula – Coeliaria – Cranophorus – Cranoryssus – Cryptolaemus – Cyrema – Cyrtaulis – Dapolia – Dioria – Empia – Epipleuria – Erithionyx – Eupalea – Eupaleoides – Exoplectra – Geodimmockius – Gordonita – Hazisia – Hoangus – Horniolus – Hovaulis – Iberorhyzobius – Incurvus – Iracilda – Keiscymnus – Leptoscymnus – Lucialla – Microrhizobius – Midus – Nat – Neorhizobius – Neoryssomus – Nephus – Nothocolus – Nurettinus – Orbipressus – Oridia – Oroides – Orynipus – Oryssomus – Paracranoryssus – Parastethorus – Peralda – Poorani – Poria – Propiptus – Pseudoazya – Pseudoryssomus – Psorolyma – Pullosidis – Rhizoryssomus – Rhyzobius – Robert – Rodatus – Roger – Sasajiscymnus – Scymniscus – Scymnodes – Scymnus – Sidonis – Siola – Stenadalia – Stethorus – Syntona – Tetrabrachys – Veronicobius – Viridigloba – Wioletta – Zagloba – Zilus

Name

Coccidulini Mulsant, 1846
Synonymy

Azyini Mulsant, 1850
Chnoodini Mulsant, 1850
Cranophorini Mulsant, 1850
Exoplectrini Crotch, 1874
Oryssomini Gordon, 1974
Poriini Mulsant, 1850
Scymnillini Casey, 1899
Scymnini Mulsant, 1846
Stethorini Dobzhansky, 1924
Tetrabrachini Kapur, 1948

References
Primary references

Mulsant, E. 1846. Histoire Naturelle des Coléoptères de France. Sulcicolles-Sécuripalpes. Paris: Maison xxiv + 26 pp. + 280 pp. + 1 pl. Full text Google Books Reference page.
Seago, A.E., Giorgi, J.A., Li, J-H. & Ślipiński, A. 2011. Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data. Molecular Phylogenetics and Evolution 60: 137–151. DOI: 10.1016/j.ympev.2011.03.015 Paywall. Full text PDF Reference page.
Nedvěd, O. & Kovář, I., 2012. Appendix: List of Genera in Tribes and Subfamilies, pp. 526-531. In: Hodek, I., van Emden, H.F. & Honěk, A. (Eds.) Ecology and Behaviour of the Ladybird Beetles (Coccinellidae), First Edition. Blackwell Publishing Ltd. 561pp. ISBN 9781405184229 DOI: 10.1002/9781118223208 Full text Reference page.

Additional references

Gordon, R.D., 2007. South American Coccinellidae (Coleoptera). Part XII (lege XIII): Carinoscymnus, new genus of South American Scymnini (Scymninae). Frustula Entomologica n.s., 30(43): 161-176. Reference page.
Tomaszewska, W. & Ślipiński, A. 2012. Oroides pakaluki new genus and new species of Coccidulini from New Guinea (Coleoptera: Coccinellidae). Zootaxa 3174: 44–50. Preview DOI: 10.11646/zootaxa.3174.1.4 ResearchGate Reference page.
González, G., 2013. Gordonita new genus and other contributions to the knowledge of South American Chnoodini (Coleoptera: Coccinellidae). Boletín Sociedad Entomológica Aragonesa, 53: 63–79. Full text PDF Reference page.
Churata-Salcedo, J.M., Almeida, L.M., González, G. & Gordon, R.D. 2017. On the taxonomy of the genus Sidonis Mulsant, stat. nov. (Coleoptera: Coccinellidae: Chnoodini) with descriptions of new species from Brazil. Zootaxa 4350(3): 500–510. DOI: 10.11646/zootaxa.4350.3.4 Full text PDF Reference page.