Boloria

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Heliconiinae
Tribus: Argynnini
Genus: Boloria
Subgenera: B. (Boloria) – B. (Clossiana) – B. (Proclossiana)
Name

Boloria Moore, 1900

Type species: Papilio pales Denis & Schiffermüller, 1775

Synonyms

Smoljana Slivov, 1995

References

Moore, A. 1900. Dinophilus gardineri (Sp. nov.). The Biological bulletin. 1: 15-18. BHL. Reference page.
Churkin, S.V., New taxa of butterflies from Transbaikalia, Russia (Lepidoptera: Satyridae; Nymphalidae). Atalanta 29 (1-4): 107–124.
Churkin, S.V., 2000: Two new subspecies of the genus Boloria Moore (Lepidoptera, Nymphalidae) from Russia.
Churkin, S.V. & Bogdanov, P.V. 2003. Two new subspecies of the genus Boloria Moore (Lepidoptera, Nymphalidae) from Siberia and Mongolia Helios 4: 217–223. Reference page.
Dubatolov V.V. & Kosterin O.E., 2010: What is Matveev's Fritillary (Clossiana matveevi P. Gorbunov et Korshunov in Korshunov, Gorbunov, 1995). Amurian zoological journal II(2): 184–188. Full article: [1].
Hsu, Y.-F. & Yen, S.-H. 1997. Notes on Boloria pales yangi, ssp. nov., a remarkable disjunction in butterfly biogeography (Lepidoptera: Nymphalidae). The Journal of Research on the Lepidoptera 34: 1–5. PDF. Reference page.
Korb S. K. & R.V. Yakovlev, 1997: Description d’un taxon nouveau de Russie: Boloria angarensis shelkovnikovi ssp. n (Lepidoptera, Nymphalidae). Alexanor 20(1): 45–46.
Perkins jr., E.M. & W.C. Meyer, 1973: Revision of the Boloria epithore complex, with description of two new subspecies (Nymphalidae). Bulletin of the Allyn Museum 11: 1–23. Full article: [2].
Simonsen, T.J. 2005. Boloria phylogeny (Lepidoptera: Nymphalidae): tentatively reconstructed on the basis of male and female genitalic morphology. Systematic Entomology 30(4): 653–665. DOI: 10.1111/j.1365-3113.2005.00292.x Paywall. Reference page.
Simonsen, T.J. 2006. Fritillary phylogeny, classification, and larval host plants: reconstructed mainly on the basis of male and female genitalic morphology (Lepidoptera: Nymphalidae: Argynnini). Biological Journal of the Linnean Society 89(4): 627–673. DOI: 10.1111/j.1095-8312.2006.00697.x Open access. Reference page.
Simonsen, T.J., Wahlberg, N., Brower, A.V.Z. & de Jong, R. 2006. Morphology, molecules and fritillaries: approaching a stable phylogeny for Argynnini (Lepidoptera: Nymphalidae). Insect Systematics & Evolution 37: 405–418. DOI: 10.1163/187631206788831407 Paywall. PDF. Reference page.
Simonsen, T.J., Wahlberg, N., Warren, A.D. & Sperling, F.A.H. 2010. The evolutionary history of Boloria (Lepidoptera: Nymphalidae): phylogeny, zoogeography and larval–foodplant relationships. Systematics and Biodiversity 8(4): 513–529. DOI: 10.1080/14772000.2010.532833 Paywall. Reference page.
Tshikolovets, V.V. 2017. Description of new taxa of Nymphalidae (Lepidoptera: Papilionoidea) from China. Zootaxa 4294(4): 480–486. DOI: 10.11646/zootaxa.4294.4.7. Reference page.

Boloria is a brush-footed butterfly (Nymphalidae) genus. Clossiana is usually included with it nowadays, though some authors still consider it distinct and it seems to warrant recognition as a subgenus at least.
Contents

1 Species
2 Brachiopod
3 References
4 Further reading
5 External links

Species

Listed alphabetically:[1][2][3]

Boloria acrocnema Gall & Sperling, 1980 – Uncompahgre fritillary
Boloria alaskensis (Holland, 1900) – mountain fritillary (Arctic America, Alaska to Hudson Bay, Wyoming, Polar Urals, Yamal Peninsula, Transbaikalia, Chukotka, Wrangel Island)
Boloria aquilonaris (Stichel, 1908) – cranberry fritillary
Boloria alberta (W.H. Edwards, 1890) – Alberta fritillary
Boloria angarensis (Erschoff, 1870) (Transbaikalia, South Siberia, Far East Yakutia, Polar Urals, Yamal Peninsula, Sayan, Tuva mountains, Amur, Ussuri, North Korea, Northeast China)
Boloria astarte (Doubleday, [1847]) – Astarte fritillary
Boloria bellona (Fabricius, 1775) – meadow fritillary
Boloria caucasica (Lederer, 1852) (Caucasus, Transcaucasia, Turkey)
Boloria chariclea (Schneider, 1794) – Arctic fritillary or purplish fritillary
Boloria dia (Linnaeus, 1767) – Weaver's fritillary or violet fritillary
Boloria distincta (Gibson, 1920) (Yukon to British Columbia, Polar Urals, Transbaikalia, Yakutia, Chukotka)
Boloria elatus (Staudinger, 1892) (Transbaikalia)
Boloria epithore (Edwards, [1864]) – Pacific fritillary
Boloria erda (Christoph, 1893) (Yakutia, Sayan, Transbaikalia, Magadan)
Boloria erubescens (Staudinger, 1901) (Tian-Shan, Ghissar-Darvaz, Pamirs-Alai)
Boloria eunomia (Esper, [1800]) – bog fritillary, ocellate bog fritillary
Boloria euphrosyne (Linnaeus, 1758) – pearl-bordered fritillary
Boloria freija (Thunberg, 1791) – Freya's fritillary, Freija fritillary or zigzag fritillary
Boloria frigga (Thunberg, 1791) – Frigga fritillary, willow bog fritillary
Boloria frigidalis Warren, 1944 (Altai, Sayan, Tuva mountains, Mongolia)
Boloria improba (Butler, 1877) – dingy fritillary
Boloria iphigenia (Graeser, 1888) (Japan, east Amur, Ussuri and northeast China)
Boloria gong (Oberthür, 1884) (Tibet, West China, North China)
Boloria graeca (Staudinger, 1870) – Balkan fritillary
Boloria jerdoni (Lang, 1868) – Jerdon's silverspot
Boloria kriemhild (Butler, 1877) – relict fritillary (Montana, Wyoming)
Boloria matveevi Gorbunov & Korshunov, 1995 (Altai)
Boloria napaea (Hoffmannsegg, 1804) – napaea fritillary or mountain fritillary
Boloria natazhati (Gibson, 1920) – cryptic fritillary or Beringian fritillary
Boloria neopales (Nakahara, 1926) (Sakhalin)
Boloria oscarus (Eversmann, 1844) (Siberia, Amur, Ussuri, Sakhalin, Yakutia)
Boloria pales (Denis & Schiffermüller, 1775) – Shepherd's fritillary
Boloria perryi (Butler, 1882) (southern Ussuri, North Korea, Amur)
Boloria polaris (Boisduval, [1828]) – Polaris fritillary
Boloria purpurea Churkin, 1999 (Barguzin Mountains in Buryatia)
Boloria selene (Denis & Schiffermüller, 1775) – small pearl-bordered fritillary or silver-bordered fritillary
Boloria selenis (Eversmann, 1837)
Boloria sipora (Moore, [1875]) (Pamirs to Alai, Tian-Shan, West Himalaya)
Boloria thore (Hübner, [1803]) – Thor's fritillary
Boloria titania (Esper, 1793) – Titania's fritillary or purple bog fritillary
Boloria tritonia (Böber, 1812) (Baikal, Amur, Ussuri)

Brachiopod

Grunt described a brachiopod genus in 1973 under the same name. Since each code of biological nomenclature allows only for one genus with the same name, the brachiopod genus is in need of renaming.
References

"Boloria Moore, 1900" at Markku Savela's Lepidoptera and Some Other Life Forms
Fritillaries, Butterflies of Canada

Jim P. Brock, Kenn Kaufman (2003) Kaufman Field Guide to Butterflies of North America. Boston: Houghton Mifflin.