Archocyrtus

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Brachycera
Infraordo: Asilomorpha
Superfamilia: Nemestrinoidea

Familia: Acroceridae
Subfamilia: †Archocyrtinae
Genus: †Archocyrtus
Species: †A. gibbosus – †A. kovalevi
Name

Archocyrtus Ussatchov, 1968: 622

Type species: Archocyrtus gibbosus Ussatchov, 1968, by original designation.
Synonyms

Juracyrtus Nartshuk, 1996: 313 [synonymized by Gillung & Winterton (2017): 327]

References
Primary references

Gillung, J.P. & Winterton, S.L. 2017. A review of fossil spider flies (Diptera: Acroceridae) with descriptions of new genera and species from Baltic Amber. Journal of Systematic Palaeontology 16(4): 325–350. DOI: 10.1080/14772019.2017.1289566 Paywall. ResearchGate Open access. Reference page.
Nartshuk, E.P. 1996. A new fossil acrocerid fly from the Jurassic beds of Kazakhstan (Diptera: Acroceridae). Zoosystematica rossica 4(2): 313–315.
Ussatchov, D.A. 1968. [New Jurassic Asilomorpha (Diptera) in Karatau]. Entomologicheskoe Obozrenie 47: 617–628 (in Russian) [English transl. in: Entomological Review 47: 378–384]. [not seen]

Additional references

Gillung, J.P. & Winterton, S.L. 2019. Evolution of fossil and living spider flies based on morphological and molecular data (Diptera, Acroceridae). Systematic Entomology 44(4): 820–841. DOI: 10.1111/syen.12358 Open access. Reference page.

Links

ION
Nomenclator Zoologicus

Archocyrtus is an extinct genus of small-headed flies in the family Acroceridae. The genus is known from compression fossils from the Late Jurassic Karabastau Formation of Kazakhstan.[1] The genus is the only member of the subfamily Archocyrtinae.[3]

A study on the holotype specimen of Archocyrtus kovalevi was published by Khramov & Lukashevich (2019). They reported evidence of an extremely long proboscis, almost twice the length of the body of the insect. They suggested that it was probably a pollinator of extinct seed plants belonging to the order Bennettitales.[4]
Species

Two species belong to the genus Archocyrtus:

†A. gibbosus Ussatchov, 1968
†A. kovalevi (Nartshuk, 1996)[2][1]

References

Gillung, Jessica P.; Winterton, Shaun L. (2017). "A review of fossil spider flies (Diptera: Acroceridae) with descriptions of new genera and species from Baltic Amber". Journal of Systematic Palaeontology. 16 (4): 325–350. doi:10.1080/14772019.2017.1289566.
Nartshuk, E. P. (1995). "A new fossil acrocerid fly from the Jurassic beds of Kazakhstan (Diptera: Acroceridae)". Zoosystematica Rossica. 4 (2): 313–315.
Gillung, Jessica P.; Winterton, Shawn L. (2019). "Evolution of fossil and living spider flies based on morphological and molecular data (Diptera, Acroceridae)". Systematic Entomology. 44 (4): 820–841. doi:10.1111/syen.12358.
Khramov, Alexander V.; Lukashevich, Elena D. (2019). "A Jurassic dipteran pollinator with an extremely long proboscis". Gondwana Research. 71: 210–215. doi:10.1016/j.gr.2019.02.004.