Anotylus

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Staphylinoidea

Familia: Staphylinidae
Subfamilia: Oxytelinae
Tribus: Oxytelini
Genus: Anotylus
Species: A. abnormalis – A. aciculatus – A. aeneotinctus – A. aethiops – A. africanus – A. alienus – A. aliiceps – A. alpicola – A. alternans – A. amicus – A. amurensis – A. andrewesi – A. andrewesides – A. anguliceps – A. antennarius – A. anticus – A. apicalis – A. apicipennis – A. applanatifrons – †A. archaicus – A. arecae – A. arellanoae – A. armatifrons – A. armifrons – A. asperiventris – A. atriceps – A. bacillus – A. bakeri – A. balbalanensis – A. barbiellinii – A. bengalensis – A. benisculptilis – A. bernhaueri – A. besucheti – A. bidentatus – A. bigemmatus – A. birmanus – A. biroi – A. bispinosus – A. bogorensis – A. borneensides – A. borneensis – A. breviceps – A. brevipennis – A. brevisculptilis – A. brincki – A. bruchi – A. brunneipennis – A. brunneus – A. bryanti – A. bubalus – A. bururianus – A. cadaverinus – A. caffer – A. cameroni – A. carinipennis – A. cavicola – A. cephalotes – A. ceylonicus – A. championi – A. chinensis – A. chinkiangensis – A. cimicoides – A. clavatus – A. clavicornis – A. clypeonitens – A. cognatus – A. comorensis – A. complanatoides – A. complanatus – A. consanguinus – A. contiguus – A. coonoor – A. coriaceus – A. cornutides – A. cornutus – A. corrugifrons – A. crassicornis – A. crebratus – A. cribriceps – A. cribrum – A. crookesi – A. cuernavacanus – A. currus – A. curticornis – A. curtipennis – A. darwini – A. darwinianus – A. debae – A. deboeri – A. deceptor – A. decipiens – A. declivisculptilis – A. delicatus – A. delopsoideus – A. densus – A. denticollis – A. dentifer – A. dentifrons – A. dilutipennis – A. discipennis – A. dispar – A. disparatus – A. distincticollis – A. drescheri – A. ebonus – A. elephantis – A. emeritus – A. engeli – A. erythrinus – A. exaratus – A. exasperatus – A. excisicollis – A. exiguus – A. extensicornis – A. externus – A. extrasculptilis – A. fageli – A. fairmairei – A. falsus – A. flavicornis – A. flavior – A. flavipennis – A. forsteri – A. fortesculpturatus – A. fortipennis – A. foveicollis – A. fragilis – A. franzi – A. frater – A. fraterculus – A. fraternus – A. frugicola – A. funebris – A. fusciventris – A. ganapati – A. garambanus – A. gardneri – A. gedyei – A. genalis – A. gibbulus – A. glaber – A. glareosus – A. gracilicornis – A. granadillae – A. grandiceps – A. gratus – A. gregarius – A. gridellii – A. gughensis – A. gunung – A. hamatoides – A. hamatus – A. hameedi – A. hammondi – A. hamuliger – A. hartmanni – A. helcopterus – A. heterocerus – A. heterophthalmus – A. hirtulus – A. holdhausi – A. hosodai – A. hostilis – A. hybridus – A. ijen – A. impennis – A. impressifrons – A. inaequalis – A. incilis – A. inornatus – A. insecatus – A. insignitus – A. intricatus – A. inustus – A. jacobsoni – A. jaljulensis – A. jamaicensis – A. jambi – A. japonicus – A. jarrigei – A. kabasi – A. kashmiricus – A. khachikovi – A. kinangopensis – A. kokodanus – A. kraatzi – A. kraussi – A. laetus – A. laevissimus – A. laticeps – A. laticornis – A. latifrons – A. latiusculus – A. leleupi – A. lemurianus – A. lewisius – A. liliputanus – A. linaxi – A. lippensi – A. lobatus – A. loebli – A. lucidulus – A. lucidus – A. lungwensis – A. luridipennis – A. luzonicus – A. madecassus – A. magdalenae – A. magniceps – A. mahmoodi – A. malaisei – A. malayanus – A. manchuricus – A. marginatus – A. maritimus – A. marmoratus – A. mashonensis – A. meccii – A. medius – A. megacephalus – A. melas – A. mendus – A. methneri – A. micans – A. microphthalmus – A. micropterus – A. micros – A. miles – A. militaris – A. mimulus – A. minutus – A. miriceps – A. mirus – A. misellus – A. mixtus – A. modestus – A. monodon – A. montanus – A. monticola – A. mortuorum – A. munitus – A. mutator – A. myops – A. myrmecophilus – A. nanus – A. neotomae – A. nigelisculptilis – A. niger – A. nigricans – A. nigripennides – A. nitelisculptilis – A. nitescens – A. nitidifrons – A. nitiduloides – A. nitidulus – A. nitouensis – A. novaeguineae – A. obliquestriatus – A. oblongifer – A. obscuratus – A. obscurellus – A. obscurifrons – A. obscurus – A. occipitalis – A. occultus – A. ocularis – A. okahandjanus – A. opacellus – A. opaciceps – A. opacicollis – A. opacifrons – A. opacinus – A. pagsanjanensis – A. panaonensis – A. papuanus – A. pedator – A. peruvianus – A. petzi – A. piceicollis – A. picticornis – A. placusinus – A. plagiatus – A. planaticollis – A. planicollis – A. plasoni – A. plumbeus – A. pluvius – A. politulus – A. politus – A. pseudopsinus – A. pumiloides – A. pumilus – A. pusillimus – A. quadricarinatus – A. quadricollis – A. quinquesulcatus – A. raffrayi – A. rectisculptilis – A. reitteri – A. rhopalocerus – A. riedeli – A. robusticornis – A. rubeculus – A. ruber – A. rubicundus – A. rubidus – A. rufinodis – A. rufus – A. rugifrons – A. rugosoides – A. rugosus – A. rugulosus – A. rurukan – A. sanctus – A. saulcyi – A. scabrellus – A. scabrifrons – A. scabripennis – A. sceptus – A. schatzmayri – A. schawalleri – A. schillhammeri – A. schuelkei – A. scorpio – A. scotti – A. sculptiventris – A. sculpturatus – A. sculptifrons – A. semipolitus – A. semiruber – A. semirufus – A. sericeiceps – A. serus – A. seticornis – A. sexualis – A. seydeli – A. shafqati – A. sharpi – A. sikkimi – A. simlaensis – A. sobrinus – A. sordidus – A. sparsus – A. speculiceps – A. speculifrons – A. spinicornis – A. spinifer – A. spinifrons – A. spinosus – A. stanleyi – A. steineri – A. stipes – A. striatellus – A. striaticeps – A. strigicollis – A. strigifrons – A. strigosulus – A. striolicollis – A. subaeneus – A. subanophtalmicus – A. subnitidus – A. subsculpturatus – A. subsericeus – A. subtilis – A. sulciceps – A. sulcicollis – A. sulcifer – A. sumatranus – A. sumatrensis – A. suspectus – A. sutteri – A. syriacus – A. szechuanensis – A. tanator – A. tardus – A. tasneemae – A. tenuesculpturatus – A. tenuicornis – A. tenuistrigosus – A. testaceicollis – A. testaceus – A. tetracarinatus – A. tetratoma – A. tinchialitensis – A. tolaensis – A. topali – A. torretassoi – A. toxopei – A. transversipennis – A. trisulcicollis – A. tuberculatus – A. tunariensis – A. turneri – A. uncifer – A. urundianus – A. ussuricus – A. varisculptilis – A. varius – A. vegrandis – A. v-elevatus – A. viator – A. vicinus – A. vilis – A. vinsoni – A. vulcanus – A. wattsensis – A. witteanus – A. wrasei – A. zavadili

Invalid species names: A. brevipennis – A. cassagnaui – A. chapmani – A. corcyranus – A. nigripennis – A. rufotestacea – A. shibatai – A. taiwana
Name

Anotylus Thomson, 1859: 44

Type species: Oxytelus sculpturatus Gravenhorst, fixed by original designation and monotypy.

Synonymy

Styloxys Gozis, 1886: 15; Herman, 1970: 414 [synonym of Anotylus]
Type species: Staphylinus rugosus Fabricius, fixed by monotypy.
Delopsis Fauvel, 1895: 198; Blackwelder, 1952: 342 [replaced by Rimba]
Type species: Delopsis cornuta Fauvel, fixed by subsequent designation by R. Lucas, 1920: 228.
Oxytelopsis Fauvel, 1895: 199; Makranczy, 2006: 75 [synonym of Anotylus]
Type species: Oxytelopsis cimicoides Fauvel, fixed by Blackwelder, 1952: 282.
Oxytelodes Bernhauer, 1908: 290; Herman, 1970: 414 [synonym of Anotylus]
Type species: Oxytelodes holdhausi Bernhauer, fixed by monotypy.
Emopotylus Bernhauer, 1910: 359 [subgenus of Oxytelus]; Herman, 1970: 414 [synonym of Anotylus]
Type species: Oxytelus (Emopotylus) cuernavacanus Bernhauer, fixed by monotypy.
Boettcherinus Bernhauer, 1936: 82; Herman, 1970: 414 [synonym of Anotylus]
Type species: Oxytelus (Boettcherinus) planaticollis Bernhauer, fixed by subsequent designation by Blackwelder, 1952: 79.
Oncoparia Bernhauer, 1936: 214; Hammond, 1976: 145 [synonym of Anotylus]
Type species: Oncoparia parasita Bernhauer, fixed by monotypy.
Paracaccoporus Steel, 1948: 188 [subgenus of Oxytelus]; Herman, 1970: 414 [synonym of Anotylus]
Type species: Oxytelus ocularis Fauvel, fixed by original designation and monotypy.
Oxytelosus Cameron, 1950: 92; Herman, 1970: 414 [synonym of Anotylus]
Type species: Oxytelus abnormalis Cameron, fixed by virtual monotypy.
Rimba Blackwelder, 1952: 342 [replacement name for Delopsis Fauvel]; Makranczy, 2006: 75 [synonym of Anotylus]
Type species: Delopsis cornuta Fauvel, fixed by objective synonym with Delopsis.
Microxytelus Fagel, 1956: 272; Herman, 1970: 414 [synonym of Anotylus]
Type species: Oxytelus nitidifrons Wollaston, fixed by original designation and monotypy.
Oxytelops Fagel, 1956: 273; Herman, 1970: 414 [synonym of Anotylus]
Type species: Staphylinus tetracarinatus Block, fixed by original designation and monotypy.
Pseudodelopsis Fagel, 1957: 3; Herman, 1970: 414 [synonym of Anotylus]
Type species: Pseudodelopsis scotti Fagel, fixed by original designation.
Anotylops Fagel, 1957: 8; Hammond, 1976: 145 [synonym of Anotylus]
Type species: Anotylops seydeli Fagel, fixed by original designation and monotypy.
Metoxytelus Coiffait & Saiz, 1968: 422; Hammond, 1976: 145 [synonym of Anotylus]
Type species: Oxytelus sulcicollis Gemminger and Harold, fixed by original designation and monotypy.
Neoplatystethus Abdullah & Qadri, 1970: 127; Hammond, 1976: 146 [synonym of Anotylus]
Type species: Neoplatystethus hameedi Abdullah & Qadri, fixed by original designation.
Pseudoplatystethus Abdullah & Qadri, 1970: 129 [subgenus of Neoplatystethus]; Hammond, 1976: 146 [synonym of Anotylus]
Type species: Neoplatystethus (Pseudoplatystethus) meccii Abdullah & Qadri, fixed by original designation.
Neopyctocraerus Abdullah & Qadri, 1970: 126; Hammond, 1976: 146 [synonym of Anotylus]
Type species: Neopyctocraerus shafqati Abdullah & Qadri, fixed by original designation and monotypy.
Pseudopyctocraerus Abdullah & Qadri, 1970: 125 [subgenus of Platystethus]; Hammond, 1976: 146 [synonym of Anotylus]
Type species: Platystethus (Pseudopyctocraerus) mahmoodi Abdullah & Qadri, fixed by original designation.

References
Primary references

Abdullah, M. & Qadri, N.-N. 1970. The Staphylindiae, Coleoptera of Pakistan. Part III. – A key to the genera and species of the Piestinae, Osoriinae, Pseudopsinae and Oxytelinae, with descriptions of new genera, subgenera and species from Karachi. Pakistan Journal of Science and Industrial Research 13(1–2): 114–131. Abstract only seen. Reference page.
Thomson, C.G. 1859: Skandinaviens Coleoptera, synoptiskt bearbetade. Tom. I. Lund: Berlingska Boktryckeriet, [5] + 290 pp.
Hammond, P.M. 1976: A review of the genus Anotylus C.G. Thomson (Coleoptera: Staphylinidae). Bulletin of the British Museum (Natural History), entomology 33(2): 137–187. BHL Reference page. [syn. nov.: p. 146]
Herman, L.H. 1970. Phylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History 142: 343–454. Handle Reference page. [syn. nov.: p. 414]
Makranczy, G. 2006. Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). Annales historico-naturales Musei nationalis hungarici 98: 29–120. Abstract and full article (PDF) Reference page. [syn. nov.: p. 75]

Additional references

Hammond, P.M. 1976: The crassicornis species-group of the genus Anotylus (Staphylinidae). Systematic entomology 1(1): 19–51. DOI: 10.1111/j.1365-3113.1976.tb00029.x Reference page.
Makranczy, G. 2011a. Four new Neotropical species of Anotylus with an interesting sexual dimorphism (Coleoptera, Staphylinidae: Oxytelinae). Annales historico-naturales Musei nationalis hungarici 103: 43–64. Reference page.
Makranczy, G. 2013: Description of a new Anotylus species from Madagascar and the female of Anotylus flavipennis from Laos (Coleoptera: Staphylinidae: Oxytelinae). Annales historico-naturales Musei nationalis hungarici 59(3): 187–195. Reference page.
Makranczy, G. 2017. Review of the Anotylus cimicoides species group (Coleoptera: Staphylinidae: Oxytelinae). Acta Zoologica Academiae Scientiarum Hungaricae 63(2): 143–262. PDF. Reference page.
Schülke, M., 2009: Zwei neue Arten der Gattung Anotylus Thomson aus der Verwandtschaft von A. complanatus (Erichson) (Coleoptera, Staphylinidae, Oxytelinae). Linzer Biologische Beiträge, 41 (2): 2009- 2024. Full article: [1].
Wang, L-F., Zhou, H-Z. & Lü, L. 2017. Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China. Zootaxa 4351(1): 1–79. DOI: 10.11646/zootaxa.4351.1.1 Reference page.

Anotylus is a genus of spiny-legged rove beetles in the family Staphylinidae. There are more than 90 described species in Anotylus.[1][2][3]
Anotylus rugosus

Species

These 95 species belong to the genus Anotylus:

Anotylus affinis (Czwalina, 1871)
Anotylus alpicola (Casey, 1894)
Anotylus antennarius Bernhauer, 1907
Anotylus arellanoae Makranczy, 2011
Anotylus athenensis (R.Dvorak, 1954)
Anotylus benisculptilis Wang, Zhou & Lü, 2017
Anotylus bernhaueri (Ganglbauer, 1898)
Anotylus breviceps (Casey, 1894)
Anotylus brevipennis (Fauvel, 1872)
Anotylus brevisculptilis Wang, Zhou & Lü, 2017
Anotylus caffer (Erichson, 1840)
Anotylus chinkiangensis Bernhauer, 1938
Anotylus clavatus (Strand, 1946)
Anotylus clypeonitens (Pandellé, 1867)
Anotylus cognatus Sharp, 1874
Anotylus complanatus (Erichson, 1839)
Anotylus coonoor Makranczy, 2017
Anotylus corcyranus Coiffait, 1968
Anotylus debae Makranczy, 2011
Anotylus deboeri Makranczy, 2011
Anotylus densus (Casey, 1894)
Anotylus engeli Makranczy, 2011
Anotylus exasperatus (Kraatz, 1859)
Anotylus exiguus (Erichson, 1840)
Anotylus extrasculptilis Wang, Zhou & Lü, 2017
Anotylus fairmairei (Pandellé, 1867)
Anotylus fraternus (Cameron)
Anotylus ganapati Makranczy, 2017
Anotylus glareosus (Wollaston, 1854)
Anotylus gunung Makranczy, 2017
Anotylus hamatus (Fairmaire & Laboulbène, 1856)
Anotylus hamuliger (Fauvel, 1905)
Anotylus hartmanni Makranczy, 2017
Anotylus herculis (Bernhauer, 1936)
Anotylus hybridus (Eppelsheim, 1878)
Anotylus ijen Makranczy, 2017
Anotylus incilis (Sharp, 1887)
Anotylus insecatus (Gravenhorst, 1806)
Anotylus insignitus (Gravenhorst, 1806)
Anotylus intricatus (Erichson, 1840)
Anotylus inustus (Gravenhorst, 1806)
Anotylus jambi Makranczy, 2017
Anotylus kabasi Makranczy, 2017
Anotylus latiusculus (Kraatz, 1859)
Anotylus linaxi Makranczy, 2017
Anotylus luridipennis (Luze, 1904)
Anotylus maritimus Thomson, 1861
Anotylus mendus Herman, 1970
Anotylus mimulus (Sharp, 1874)
Anotylus munitus (Casey, 1894)
Anotylus mutator (Lohse, 1963)
Anotylus nanus (Erichson, 1840)
Anotylus neotomae (Hatch, 1957)
Anotylus nigelisculptilis Wang, Zhou & Lü, 2017
Anotylus niger (LeConte, 1877)
Anotylus nitelisculptilis Wang, Zhou & Lü, 2017
Anotylus nitidifrons (Wollaston, 1871)
Anotylus nitidulus (Gravenhorst, 1802)
Anotylus obscurellus (Fauvel, 1905)
Anotylus opaciceps Bernhauer, 1938
Anotylus petzi (Bernhauer, 1914)
Anotylus placusinus (LeConte, 1877)
Anotylus plagiatus (Rosenhauer, 1856)
Anotylus politus (Erichson, 1840)
Anotylus pumilus (Erichson, 1839)
Anotylus rectisculptilis Wang, Zhou & Lü, 2017
Anotylus riedeli Makranczy, 2017
Anotylus rugifrons (Hochhuth, 1849)
Anotylus rugosus (Fabricius, 1775)
Anotylus rurukan Makranczy, 2017
Anotylus saulcyi (Pandellé, 1867)
Anotylus schatzmayri (Koch, 1937)
Anotylus schawalleri Makranczy, 2017
Anotylus schillhammeri Makranczy, 2017
Anotylus schuelkei Makranczy, 2017
Anotylus sculpturatus (Gravenhorst, 1806)
Anotylus sikkimi Fauvel, 1905
Anotylus sobrinus (LeConte, 1877)
Anotylus spec Makranczy, 2017
Anotylus speculifrons (Kraatz, 1857)
Anotylus steineri Makranczy, 2013
Anotylus subanophthalmicus Outerelo, Gamara & Salgado, 1998
Anotylus subsericeus Bernhauer, 1938
Anotylus suspectus (Casey, 1894)
Anotylus tanator Makranczy, 2017
Anotylus tetracarinatus (Block, 1799)
Anotylus tetratoma (Czwalina, 1871)
Anotylus topali Makranczy, 2017
Anotylus torretassoi (Koch, 1932)
Anotylus varisculptilis Wang, Zhou & Lü, 2017
Anotylus vegrandis (Casey, 1894)
Anotylus vicinus Sharp, 1874
Anotylus vinsoni (Cameron, 1936)
Anotylus zavadili (Roubal, 1941)
† Oxytelus gibbulus (Eppelsheim, 1877)

References

"Anotylus Report". Integrated Taxonomic Information System. Retrieved 2019-09-23.
"Anotylus". GBIF. Retrieved 2019-09-23.

"Anotylus genus Information". BugGuide.net. Retrieved 2019-09-23.