Gnaphosidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Chelicerata
Classis: Arachnida
Ordo: Araneae
Subordo: Opisthothelae
Infraordo: Araneomorphae
Taxon: Neocribellatae
Series: Entelegynae
Sectio: Dionycha
Superfamilia: Gnaphosoidea

Familia: Gnaphosidae
Subfamiliae: Drassodinae - Echeminae - Encoptarthriinae - Gnaphosinae - Herpyllinae - Laroniinae - Leptodrassinae - Micariinae - Prodidominae - Solitudinae - Zelotinae - †Eomactatorinae
Overview of genera (118)

Allomicythus – Allozelotes – Almafuerte – Amazoromus – Amusia – Anagraphis – Aneplasa – Anzacia – Aphantaulax – Apodrassodes – Apodrassus – Apopyllus – Aracus – Arauchemus – Arboricaria – Asemesthes – Asiabadus – Australoechemus – Battalus – Benoitodes – Berinda – Berlandina – Cabanadrassus – Callilepis – Callipelis – Camillina – Ceryerda – Cesonia – Cladothela – Coillina – Coreodrassus – Cryptodrassus – Cubanopyllus – Diaphractus – Drassodes – Drassodex – Drassyllus – Echemella – Echemographis – Echemoides – Echemus – Eilica – Encoptarthria – Epicharitus – Fedotovia – Gaviphosa – Gertschosa – Gnaphosa – Haplodrassus – Hemicloea – Herpyllus – Heser – Hitobia – Homoeothele – Hongkongia – Hotwheels – Hypodrassodes – Intruda – Iranotricha – Kaitawa – Kishidaia – Ladissa – Laronius – Latonigena – Leptodrassex – Leptodrassus – Leptopilos – Litopyllus – Marinarozelotes – Marusik (Lin & Li) – Matua – Megamyrmaekion – Meizhelan – Micaria – Microdrassus – Microsa – Micythus – Minosia – Minosiella – Montebello – Nauhea – Nodocion – Nomisia – Notiodrassus – Odontodrassus – Orodrassus – Parabonna – Parasyrisca – Phaeocedus – Platnickus – Poecilochroa – Pseudodrassus – Pterotricha – Pterotrichina – Sanitubius – Scopoides – Scotocesonia – Scotognapha – Scotophaeus – Sergiolus – Sernokorba – Setaphis – Shaitan – Shiragaia – Sidydrassus – Smionia – Sosticus – Symphanodes – Synaphosus – Talanites – Talanitoides – Titus – Trachyzelotes – Trephopoda – Trichothyse – Urozelotes – Vectius – Xenoplectus – Xerophaeus – Xizangia – Yuqilin – Zelanda – Zelominor – Zelotes – Zelotibia – Zimiromus – †Eognaphosops – †Eomactator – †Zelotetis
Name

Gnaphosidae Pocock, 1898
Synonyms

Drassidae Sundevall, 1833

References
Primary references

Chatzaki, M. & Russell-Smith, A. 2017. New species and new records of ground spiders (Araneae: Gnaphosidae) from Cyprus. Zootaxa 4329(3): 237–255. DOI: 10.11646/zootaxa.4329.3.3. Reference page.
Fomichev, A. & Marusik, Y.M. 2015. A survey of East Palaearctic Gnaphosidae (Araneae). 4. A review of Fedotovia Charitonov, 1946. Zootaxa 3948(1): 93–108. DOI: 10.11646/zootaxa.3948.1.6. Preview (PDF). Reference page.
Grismado, C.J. & López Carrión, N. 2017. Description of Almafuerte, a new genus of ground spiders from South America (Araneae, Gnaphosidae). Zootaxa 4338(2): 263–291. DOI: 10.11646/zootaxa.4338.2.3. Reference page.
Lin, Y-J. & Li, S. 2023a. On nine ground spiders from Xishuangbanna, China (Araneae, Gnaphosidae), including two new genera and seven new species. Zookeys 1174ː 141–174. DOI: 10.3897/zookeys.1174.106340 Open access Reference page.
Pocock, R.I. 1898. The Arachnida from the province of Natal, South Africa, contained in the collection of the British Museum. Annals and Magazine of Natural History (7)2: 197–226. [219]
Zamani, A., Seiedy, M., Saboori, A. & Maruski, Yu.M. 2018. The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae). ZooKeys 777: 17–41. DOI: 10.3897/zookeys.777.26745 Reference page.

Additional references

Bosmans, R. 2014: On the identity of the genera Anagraphis Simon, 1893 and Macedoniella Drensky, 1935 with two new synonyms (Araneae: Gnaphosidae). Arachnologische Mitteilungen 48: 38–41. full article (PDF). Reference page.
Liu, B. & Zhang, F. 2023a. Revision of the genus Scopoides Platnick, 1989 from China, with description of a new genus (Araneae, Gnaphosidae). Zookeys 1172ː 203–215. DOI: 10.3897/zookeys.1172.105034 Open access Reference page.
Liu, B. & Zhang, F. 2024. Hotwheels gen. nov., a new ground spider genus (Araneae, Gnaphosidae) from southwest China. ZooKeys 1189: 337–347. DOI: 10.3897/zookeys.1189.115996 Open access Reference page.
Murphy, J. 2007. Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. ISBN 978 0 9500093 4 6 flier [not seen]

Vernacular names
Deutsch: Plattbauchspinnen
English: Ground spider
日本語: ワシグモ科
한국어: 수리거미과
Nederlands: Bodemjachtspinnen
中文: 平腹蛛科

Ground spiders comprise Gnaphosidae, the seventh largest spider family with over 2,000 described species in over 100 genera distributed worldwide. There are 105 species known to central Europe,[1] and common genera include Gnaphosa, Drassodes, Micaria, Cesonia, Zelotes and many others. They are closely related to Clubionidae.[2] At present, no ground spiders are known to be seriously venomous to humans.
Description

Generally, ground spiders are characterized by having barrel-shaped anterior spinnerets that are one spinneret diameter apart. The main exception to this rule is found in the ant-mimicking genus Micaria. Another characteristic is an indentation in the endites (paired mouthparts anterior and lateral to the labium, or lip). All ground spiders lack a prey-capture web and generally run prey down on the surface. They hunt at night and spend the day in a silken retreat.[2] The genitalia are diverse and are a good model for studying the evolution of genitalia because of their peculiar copulatory mechanism.[3] The thick-walled egg sacs are guarded by the mother until the spiderlings hatch.[2]
Hunting behavior

Ground spiders hunt by active foraging, chasing down and subduing individual prey items. They are adapted to hunting large and potentially dangerous prey, including other spiders, which they subdue by using their silk. When hunting, ground spiders produce thick, gluey silk from their enlarged spinnerets and attempt to use it to entangle their prey in swathing attacks, often applying their webbing to their prey's legs and mouths. By immobilizing potential prey in this manner, ground spiders can subdue proportionally large creatures while reducing risk of injury to themselves from their prey's attempts to fight back.[4]
Genera
Main article: List of Gnaphosidae species

As of January 2024, the World Spider Catalog accepted the following genera:[5]

Allomicythus Ono, 2009 — Vietnam
Allozelotes Yin & Peng, 1998 — China
Almafuerte Grismado & Carrión, 2017 — Bolivia, Argentina, Uruguay
Amazoromus Brescovit & Höfer, 1994 — Brazil
Ammoxenus Simon, 1893
Amusia Tullgren, 1910 — South Africa, East Africa
Anagraphis Simon, 1893 — Africa, Asia, Europe
Anagrina Berland, 1920 — Niger
Aneplasa Tucker, 1923 — South Africa, Angola
Anzacia Dalmas, 1919 — Australia, New Zealand, Papua New Guinea
Aphantaulax Simon, 1878 — Asia, Africa, Australia, France
Apodrassodes Vellard, 1924 — South America, Mexico, India
Apodrassus Chamberlin, 1916 — Peru
Aponetius Kamura, 2020 — Ryukyu Is.
Apopyllus Platnick & Shadab, 1984 — South America, Mexico
Arauchemus Ott & Brescovit, 2012 — Brazil
Asemesthes Simon, 1887 — Africa
Asiabadus Roewer, 1961 — Afghanistan
Australoechemus Schmidt & Piepho, 1994 — Cape Verde Is.
Austrammo Platnick, 2002
Avstroneulanda Zakharov & Ovtsharenko, 2022
Barrowammo Platnick, 2002
Benoitodes Platnick, 1993 — St. Helena
Berinda Roewer, 1928 — Greece, Turkey, Cyprus
Berlandina Dalmas, 1922 — Asia, Africa, Europe
Cabanadrassus Mello-Leitão, 1941 — Argentina
Callilepis Westring, 1874 — Asia, North America, Europe
Callipelis Zamani & Marusik, 2017 — Iran
Camillina Berland, 1919 — Asia, Central America, South America, Africa, North America, Italy, Caribbean
Canariognapha Wunderlich, 2011 — Canary Is.
Ceryerda Simon, 1909 — Australia
Cesonia Simon, 1893 — Greece, Turkey, North America, Central America, Caribbean
Chatzakia Lissner & Bosmans, 2016 — Spain
Civizelotes Senglet, 2012 — Asia, Morocco, Europe
Cladothela Kishida, 1928 — Japan, China, Korea
Coillina Yin & Peng, 1998 — China
Coreodrassus Paik, 1984 — Asia
Cryptodrassus Miller, 1943 — Europe, Asia
Cryptoerithus Rainbow, 1915 — Australia
Cubanopyllus Alayón & Platnick, 1993 — Cuba
Diaphractus Purcell, 1907 — South Africa, Namibia
Drassodes Westring, 1851 — Africa, South America, Asia, Europe, North America
Drassodex Murphy, 2007 — Spain, France, Switzerland
Drassyllus Chamberlin, 1922 — North America, Asia, Europe
Echemella Strand, 1906 — Congo, Ethiopia
Echemographis Caporiacco, 1955 — Venezuela
Echemoides Mello-Leitão, 1938 — South America
Echemus Simon, 1878 — Sweden, Asia, Australia, Africa, Brazil
Eilica Keyserling, 1891 — Australia, South America, United States, Central America, Caribbean, Africa, India
Encoptarthria Main, 1954 — Australia
Epicharitus Rainbow, 1916 — Australia
Fedotovia Charitonov, 1946 — Mongolia, Iran, Afghanistan
Gaviphosa Sankaran, 2021 — India
Gertschosa Platnick & Shadab, 1981 — North America, Panama, Jamaica
Gnaphosa Latreille, 1804 — Asia, Europe, Africa, North America, Guatemala, Cuba
Haplodrassus Chamberlin, 1922 — Europe, Asia, North America, Africa
Herpyllus Hentz, 1832 — Argentina, North America, Asia
Heser Tuneva, 2004 — Asia, Spain, Africa, North America
Hitobia Kamura, 1992 — Asia
Homoeothele Simon, 1908 — Australia
Hotwheels Liu & Zhang, 2024 — China
Hongkongia Song & Zhu, 1998 — Indonesia, China, Hong Kong
Hypodrassodes Dalmas, 1919 — New Zealand, Australia
Ibala Fitzpatrick, 2009 — Africa
Intruda Forster, 1979 — Australia, New Zealand
Iranotricha Zamani & Marusik, 2018 — Iran
Kaitawa Forster, 1979 — New Zealand
Kikongo Rodrigues & Rheims, 2020 — D.R. Congo, Kenya
Kishidaia Yaginuma, 1960 — Asia, Italy
Kituba Rodrigues & Rheims, 2020 — D.R. Congo
Ladissa Simon, 1907 — Sierra Leone, India, Benin
Laronius Platnick & Deeleman-Reinhold, 2001 — Thailand, Indonesia
Lasophorus Chatzaki, 2018 — Greece
Latica da Silva, Guerrero, Bidegaray-Batista & Simó, 2020 — Uruguay, Argentina
Latonigena Simon, 1893 — South America
Leptodrassex Murphy, 2007 — Africa, Europe, Asia
Leptodrassus Simon, 1878 — Europe, Asia, Africa, Mexico
Leptopilos Levy, 2009 — Greece, Asia, Libya
Litopyllus Chamberlin, 1922 — United States, Cuba, Mexico
Macarophaeus Wunderlich, 2011 — Canary Is., Madeira
Marinarozelotes Ponomarev, 2020 — Asia, Europe, Africa, North America, South America
Marjanus Chatzaki, 2018
Matua Forster, 1979 — New Zealand
Megamyrmaekion Reuss, 1834 — Africa, Asia, Australia
Micaria Westring, 1851 — Asia, North America, Africa, Europe, Australia
Microdrassus Dalmas, 1919 — Seychelles
Microsa Platnick & Shadab, 1977 — Cuba
Micythus Thorell, 1897 — Thailand, Myanmar, Indonesia
Minosia Dalmas, 1921 — Africa, Asia, Europe
Minosiella Dalmas, 1921 — Asia, Africa
Molycria Simon, 1887 — Australia
Montebello Hogg, 1914 — Australia
Myandra Simon, 1887 — Australia
Nauhea Forster, 1979 — New Zealand
Neodrassex Ott, 2012 — Brazil, Argentina
Nodocion Chamberlin, 1922 — North America, India
Nomindra Platnick & Baehr, 2006 — Australia
Nomisia Dalmas, 1921 — Asia, Africa, Europe
Notiodrassus Bryant, 1935 — New Zealand
Odontodrassus Jézéquel, 1965 — Africa, Asia, Jamaica
Orodrassus Chamberlin, 1922 — United States, Canada
Parabonna Mello-Leitão, 1947 — Brazil
Parasyrisca Schenkel, 1963 — Europe, Asia, North America
Phaeocedus Simon, 1893 — Asia, Europe
Poecilochroa Westring, 1874 — Asia, Africa, South America, Europe
Pseudodrassus Caporiacco, 1935 — China, Libya, Turkey
Pterotricha Kulczyński, 1903 — Africa, Asia, Europe
Pterotrichina Dalmas, 1921 — Algeria
Rastellus Platnick & Griffin, 1990
Sanitubius Kamura, 2001 — China, Korea, Japan
Scopoides Platnick, 1989 — North America, Asia
Scotocesonia Caporiacco, 1947 — Guyana
Scotognapha Dalmas, 1920 — Canary Islands
Scotophaeus Simon, 1893 — Europe, Africa, Asia, South America, New Zealand
Sergiolus Simon, 1892 — North America, Caribbean, Asia
Sernokorba Kamura, 1992 — Asia, Europe
Setaphis Simon, 1893 — Spain, Africa, Asia
Shaitan Kovblyuk, Kastrygina & Marusik, 2013
Shiragaia Paik, 1992 — Korea
Sidydrassus Esyunin & Tuneva, 2002 — Asia
Smionia Dalmas, 1920 — South Africa
Solitudes Lin & Li, 2020 — China
Sosticus Chamberlin, 1922 — North America, Asia
Symphanodes Rainbow, 1916 — Australia
Synaphosus Platnick & Shadab, 1980 — Asia, Africa, North America, Greece
Talanites Simon, 1893 — Asia, North America, Ukraine
Talanitoides Levy, 2009 — Israel
Titus O. Pickard-Cambridge, 1901 — Zimbabwe
Trachyzelotes Lohmander, 1944 — Europe, Asia, North America, Africa, South America, Samoa
Trephopoda Tucker, 1923 — South Africa, Namibia
Trichothyse Tucker, 1923 — South Africa, Namibia
Turkozelotes Kovblyuk & Seyyar, 2009 — Greece, Asia
Urozelotes Mello-Leitão, 1938 — Zambia, Europe, Asia
Verita Ramírez & Grismado, 2016 — Argentina
Wesmaldra Platnick & Baehr, 2006 — Australia
Wydundra Platnick & Baehr, 2006 — Australia, Malaysia
Xerophaeus Purcell, 1907 — Africa, Yemen
Xizangiana Song, Zhu & Zhang, 2004 — China
Yoruba Rodrigues & Rheims, 2020 — Nigeria, Ivory Coast
Zagrotes Zamani, Chatzaki, Esyunin & Marusik, 2021 — Iran
Zelanda Özdikmen, 2009 — New Zealand
Zelominor Snazell & Murphy, 1997 — Portugal, Spain, Algeria
Zelotes Gistel, 1848 — Asia, North America, Europe, Africa, South America, Australia, Caribbean
Zelotibia Russell-Smith & Murphy, 2005 — Africa
Zelowan Murphy & Russell-Smith, 2010 — Congo, Burundi, Namibia
Zimiromus Banks, 1914 — Central America, South America, Caribbean, Mexico