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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Lamiales

Familia: Plantaginaceae
Tribus: Gratioleae
Genus: Gratiola
Species: G. amphiantha – G. aurea – G. brevifolia – G. ebracteata – G. flava – G. floridana – G. fluviatilis – G. graniticola – G. griffithii – G. heterosepala – G. hispida – G. japonica – G. linifolia – G. mauretanica – G. nana – G. neglecta – G. officinalis – G. oresbia – G. pedunculata – G. peruviana – G. pilosa – G. pumilio – G. quartermaniae – G. ramosa – G. virginiana – G. viscidula
Name

Gratiola L., Sp. Pl. 1: 17. (1753)

Type species: Gratiola officinalis L., Sp. Pl. 1: 17. (1753)

Synonyms

Heterotypic
Amphianthus Torr., Ann. Lyceum Nat. Hist. New York 4: 82. (1837)

References

Linnaeus, C. 1753. Species Plantarum 1: 17.
Estes, D. & Small, R.L. 2008. Phylogenetic Relationships of the Monotypic Genus Amphianthus (Plantaginaceae Tribe Gratioleae) Inferred from Chloroplast DNA Sequences. Systematic Botany 33(1): 176–182. DOI: 10.1600/036364408783887375 JSTOR PDF Reference page.
Hassler, M. 2018. Gratiola. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2018. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2018 Nov. 7. Reference page.
International Plant Names Index. 2018. Gratiola. Published online. Accessed: Nov. 7 2018.
Tropicos.org 2018. Gratiola. Missouri Botanical Garden. Published on the internet. Accessed: 2018 Nov. 7.

Vernacular names
العربية: غراتيولا
беларуская: Аўран
català: Gracíola
čeština: konitrud
dansk: Nådesurt-slægten
Deutsch: Gnadenkräuter
English: Hedge Hyssops
فارسی: زیباساز
suomi: Kuntiot
hornjoserbsce: Hnadnička
lietuvių: Raitinytė
русский: Авран
slovenčina: graciola
svenska: Jordgallor
українська: Авран
oʻzbekcha/ўзбекча: Avran
中文: 水八角属

Gratiola is a genus of plants in the family Plantaginaceae. Most species are known generally as hedgehyssops. The genus was previously included in the family Scrophulariaceae.

Species include:

Gratiola amphiantha - pool sprite, snorkelwort
Gratiola aurea - golden hedgehyssop
Gratiola brevifolia - sticky hedgehyssop
Gratiola ebracteata - bractless hedgehyssop
Gratiola flava - yellow hedgehyssop
Gratiola floridana - Florida hedgehyssop
Gratiola heterosepala - Boggs Lake hedgehyssop
Gratiola neglecta - clammy hedgehyssop
Gratiola officinalis - common hedgehyssop
Gratiola peruviana - Austral brooklime (native to South America and Australasia)
Gratiola pubescens - Hairy brooklime (native to Australia)
Gratiola quartermaniae - limestone hedgehyssop
Gratiola ramosa - branched hedgehyssop
Gratiola virginiana - roundfruit hedgehyssop
Gratiola viscidula - Short's hedgehyssop

Fossil record

Four fossil seeds of †Gratiola tertiaria have been extracted from borehole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland.[1]
References

Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) [Szczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska)]. Acta Palaeobotanica 1979 20 (1): 3-117.

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