Falco peregrinus , Photo: Michael Lahanas
Falco peregrinus Tunstall, 1771
Falco peregrinus (*)
* Ornithologia britannica p.1
The Peregrine Falcon (Falco peregrinus), also known as the Peregrine, and historically as the "Duck Hawk" in North America, is a cosmopolitan bird of prey in the family Falconidae. The Peregrine is renowned for its speed, reaching speeds of over 320 km/h (200 mph) during its characteristic hunting stoop, making it the fastest extant member of the animal kingdom.
It is a large, crow-sized falcon, with a blue-gray back, barred white underparts, and a black head and "moustache". Experts recognize 17 to 19 subspecies which vary in appearance and range; there is disagreement over whether the distinctive Barbary Falcon is a subspecies or a distinct species. Typical of bird-eating raptors, all subspecies of F. peregrinus, including the nominate, are sexually dimorphic, with females being considerably larger than males. 
The Peregrine's breeding range includes land regions from the Arctic tundra to the Tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This fact makes it the world's most widespread bird of prey. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations.
While its diet consists almost exclusively of medium-sized birds, the Peregrine will occasionally hunt small mammals, small reptiles or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The Peregrine Falcon became an endangered species in many areas due to the use of pesticides, especially DDT. Since the ban on DDT from the beginning of the 1970s onwards, the populations recovered, supported by large scale protection of nesting places and releases to the wild.
The Peregrine Falcon has a body length of 34 to 58 centimetres (13–23 in) and a wingspan of around 80 to 120 centimetres (31–47 in). The male and female have similar markings and plumage, but as in many birds of prey the Peregrine Falcon displays marked reverse sexual dimorphism in size, with the female measuring up to 30 percent larger than the male. Males weigh 440 to 750 grams (0.97–1.7 lb) and the noticeably larger females weigh 910 to 1,500 grams (2.0–3.3 lb); for variation in weight between subspecies, see under that section below.
The back and the long pointed wings of the adult are usually bluish black to slate gray with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, colored like the back but with thin clean bars, is long, narrow and rounded at the end with a black tip and a white band at the very end. The top of the head and a "mustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. The immature bird is much browner with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.
This species was first described by Marmaduke Tunstall in his 1771 Ornithologia Britannica under its current binomial name. The scientific name Falco peregrinus, means "wandering falcon" in Latin. Indeed, the species' common name refers to its wide-ranging flights in most European languages. The Latin term for falcon, falco, is related to falx, the Latin word meaning sickle, in reference to the silhouette of the falcon's long, pointed wings in flight.
The Peregrine Falcon belongs to a genus whose lineage includes the hierofalcons and the Prairie Falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Early Pliocene, about 8–5 million years ago (mya). As the Peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear; the issue is complicated by widespread hybridization confounding mtDNA sequence analyses; for example a genetic lineage of the Saker Falcon (F. cherrug) is known which originated from a male Saker producing fertile young with a female Peregrine ancestor.
Today, Peregrines are regularly hybridized in captivity with other species such as the Lanner Falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the Peregrine's hunting skill with the Lanner's hardiness, or the Gyrfalcon to produce large, strikingly colored birds for the use of falconers. As can be seen, the Peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).
Numerous subspecies of the Peregrine have been described, with 19 accepted by the Handbook of the Birds of the World.
* Falco peregrinus peregrinus, the nominate subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 grams (1.3–1.7 lb), while females weigh 925 to 1,300 grams (2.04–2.9 lb). It includes brevirostris, germanicus, rhenanus, and riphaeus.
* Falco peregrinus calidus, described by John Latham in 1790, was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia, from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory, and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than peregrinus, especially on the crown. Males weigh 588 to 740 grams (1.30–1.6 lb), while females weigh 925 to 1,333 grams (2.04–2.94 lb).
* Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti and pleskei, and harterti seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on coast there), and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to peregrinus, but the young are even darker than those of anatum.
Australian race F. p. macropus
* Falco peregrinus macropus, described by Swainson in 1837 is the Australian Peregrine Falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.
* Falco peregrinus submelanogenys described by Mathews in 1912, is the Southwest Australian Peregrine Falcon. It is found in southwest Australia and is non-migratory.
* Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian Peregrine Falcon, Black Shaheen, or Indian Shaheen. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from Pakistan across India and Bangladesh to Sri Lanka and Southeastern China. In India, the Shaheen is reported from all states except Uttar Pradesh mainly from rocky and hilly regions. The Shaheen is also reported from the Andaman and Nicobar Islands. They have a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on manmade structures such as buildings and cellphone transmission towers. A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory, and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills while the migrant calidus is more often seen along the coast. In Pakistan it is a military symbol of the Pakistan Air Force.
* Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American Peregrine Falcon, or "Duck Hawk"; its scientific name means "Duck Peregrine Falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains today. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts seek to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to peregrinus but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 570 grams (1.1–1.3 lb), while females weigh 900 to 960 grams (2.0–2.1 lb).
* Falco peregrinus cassini, described by Sharpe in 1873, is also known as the Austral Peregrine Falcon. It includes kreyenborgi, the Pallid Falcon a leucistic morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and Falkland Islands. It is non-migratory. It is similar to nominate, but slightly smaller with a black ear region. The variation kreyenborgi is medium grey above, has little barring below, and has a head pattern like the Saker Falcon, but the ear region is white.
Captive bird of the subspecies pealei
* Falco peregrinus pealei, described by Ridgway in 1873, is also known as Peale's Falcon, and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from the Puget Sound along the British Columbia coast (including the Queen Charlotte Islands), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia. It is possibly found on the Kuril Islands and the coasts of Kamchatka as well. It is non-migratory. It is the largest subspecies, and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns.
* Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys It is found in the Arctic tundra of North America to Greenland. It migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously united with anatum. It is the New World equivalent to calidus. It is smaller than anatum. It is also paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner, and less grey, than in calidus, and paler, sometimes almost sandy, than in anatum.
* Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary Falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands, and is non-migratory; it is endangered with only six to eight pairs surviving. Males have a rufous wash on crown, nape, ears and back; underside conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.
* Falco peregrinus minor was first described by Bonaparte in 1850. It was formerly often perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory, and small and dark.
* Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and Comoros. It is non-migratory.
* Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean Peregrine Falcon or the Maltese Falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides, Barbary Falcons (see also below), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies, and the underside usually has rusty hue. Males weigh around 445 grams (0.98 lb), while females weigh up to 920 grams (2.0 lb).
Painting of subspecies babylonicus by John Gould
* Falco peregrinus ernesti, described by Sharpe in 1894, is found from Indonesia to Philippines and south to Papua New Guinea and Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate in the very dark, dense barring on its underside and its black ear coverts.
* Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands. It is non-migratory. It is very rare, and may only remain on a single island. It is a dark form, resembling pealei in color, but darker, especially on tail.
* Falco peregrinus nesiotes described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.
* Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through north Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 grams (1.3 lb).
* Falco peregrinus babylonicus described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and Tian Shan to Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides, and somewhat similar to a small, pale Lanner Falcon. Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).
These last two races are often split as Barbary Falcon Falco pelegrinoides. There is a 0.6–0.7% genetic distance in the Peregine-Barbary Falcon ("peregrinoid") complex. These birds inhabit arid regions from the Canary Islands along the rim of the Sahara through the Middle East to Central Asia and Mongolia. They have a red neck patch but otherwise differ in appearance from the Peregrine proper merely according to Gloger's Rule. The Barbary Falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the Peregrine, but less often and far less pronounced. The Barbary Falcon's shoulder and pelvis bones are stout by comparison with the Peregrine, and its feet are smaller. They have no postzygotic reproduction barriers in place, but they breed at different times of year than neighboring Peregrine Falcon subspecies.
Ecology and behavior
The life span in the wild is up to 15.5 years. Mortality in the first year is between 59–70%, declining to between 25–32% in adults. Apart from anthropogenic threats like collision with human-made objects, the Peregrine may be killed by large eagles or large owls. The Peregrine Falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the Peregrine Falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known Peregrine Falcon ectoparasites are chewing lice Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).
The Peregrine Falcon hunts at dawn and dusk, when prey are most active, but in cities also nocturnally, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by Peregrines include species as diverse as Yellow-billed Cuckoo, Black-necked Grebe, Virginia Rail and Common Quail. It requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields and tundra. It searches for prey either from a high perch or from the air. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. The air pressure from a 200 mph (320 km/h) dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. Prey is struck and captured in mid-air; the Peregrine Falcon strikes its prey with a clenched foot, stunning or killing it, then turns to catch it in mid-air. The Peregrine will drop it to the ground and eat it there if it is too heavy to carry. Prey is plucked before consumption.
The Peregrine Falcon nests in a scrape, normally on cliff edges or, today regularly in many parts of its range, on tall buildings or bridges. Cliff nests are generally located under an overhang, on ledges with vegetation, and south-facing sites are favored. In some regions, as in parts of Australia and on the west coast of Northern North-America, large tree hollows are used for nesting. Before the demise of most European peregrines, there was a large population of peregrines in central and western Europe using the disused nests of other large birds. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. The human-made structures used for breeding closely resemble the natural cliff ledges that the Peregrine prefers for its nesting locations.
Mostly three to four eggs (range 1–5) are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female. The male also helps with the incubation of the eggs over day, but at night only the female incubates. The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere (the Australian subspecies macropus may breed as late as November and equatorial populations may nest anytime between June and December). The female generally lays another clutch if the eggs are lost early in the nesting season, though this is extremely rare in the Arctic owing to the short summer season. As a result of some infertile eggs and natural losses of nestlings, the average number of young found in nests is 2.5, and the average number that fledges is about 1.5.
After hatching, the eyases, or chicks, are covered with creamy-white down and have disproportionately large feet. The male, which is called the "tiercel", and the female, which is simply called the "falcon", both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12–15 miles) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.
Relationship with humans
Peregrine eggs and chicks are often targeted by black marketeers and unscrupulous egg collectors, so it is normal practice not to publicize unprotected nest locations.
In the USA, Canada, Germany and Poland, Wildlife services in Peregrine Falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a Peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment. Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The Peregrine Falcon was removed from the U.S. Endangered Species list on August 25, 1999.
In the USA
Many Peregrine Falcons have settled in large cities, nesting on cathedrals, skyscraper window ledges, and the towers of suspension bridges. As early as 1946, Peregrine Falcons were nesting atop Philadelphia City Hall, which is believed to be among the first artificial structures in the world to be used as a nest site by this species. In Virginia, state officials working with students from the Center for Conservation Biology of the College of William and Mary in Williamsburg successfully established nesting boxes high atop the George P. Coleman Memorial Bridge on the York River, the Benjamin Harrison Memorial Bridge and Varina-Enon Bridge on the James River, and at other similar locations. Thirteen new chicks were hatched in this Virginia program during a recent year. Over 250 falcons have been released through the Virginia program. The New York State Department of Environmental Conservation reported that there were 67 pairs of Peregrine Falcons in the state during 2008. San Francisco enjoys two nesting falcons at the Pacific Gas & Electric building.
As in the USA, Peregrine Falcons have moved into major Canadian cities.
In Winnipeg, Manitoba, the first post-population crash wild nest occurred in 1989 on the Delta Winnipeg Hotel (now the Radisson Hotel Downtown-Skyview) as a result of recovery efforts begun in 1980–1981 by the Peregrine Falcon Recovery Project (Manitoba). The Radisson site has been occupied continuously since 1989 and offspring from this nest can be found across the Canadian Prairie Provinces and the US Midwest States. The popular CBC Manitoba/Shaw FalconCam has been following the lives (and deaths) of the Radisson peregrines since 2006. In 1993, peregrines began nesting at the McKenzie Seeds Building in Brandon, Manitoba and have likewise had a similarly long and successful nesting history.
In Hamilton, there have been a pair of birds nesting on a ledge overhanging a window of the downtown Sheraton hotel every year since 1994. The Hamilton Community Peregrine Project was setup in 1995 to observe the falcons and provide care should they need it. This includes banding birds that have successfully fledged and caring for birds which crash onto the busy streets below the nest. They also have a camera aimed at the nesting site, which can be viewed on their website.
The University of Alberta in Edmonton, Alberta also houses a pair of breeding Peregrine Falcons atop the Clinical Sciences building, amongst others on the campus who are actively monitored via webcams. The female of this pair originated in Winnipeg, Manitoba while the male is from Alberta. Several birds have also been tagged with radio transmitters, permitting study of migration patterns.
In Britain, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalizing on the urban pigeon populations for food. One notable urban example is the pair of breeding Peregrines that nest on a ledge on the tower of Derby Cathedral.
* A Peregrine Falcon is prominently featured on the Idaho quarter issued in 2007 as part of the United States Mint's 50 State Quarters program.
1. ^ BirdLife International (2009). Falco peregrinus. In: IUCN 2009. IUCN Red List of Threatened Species. Downloaded on 2009-11-03.
* American Ornithologists' Union (1910): Check-list of North American Birds (Third Edition) American Ornithologists' Union.
Source: Wikipedia, Wikispecies: All text is available under the terms of the GNU Free Documentation License