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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Cladus: Telluraves
Cladus: Australaves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Corvida
Superfamilia: Corvoidea

Familia: Corvidae
Genus: Cyanocitta
Species: Cyanocitta stelleri
Subspecies: C. s. annectens – C. s. azteca – C. s. carbonacea – C. s. carlottae – C. s. coronata – C. s. diademata – C. s. frontalis – C. s. macrolopha – C. s. philippsi – C. s. purpurea – C. s. restricta – C. s. stelleri – C. s. suavis
Name

Cyanocitta stelleri (Gmelin, 1788)
Synonymy

Corvus stelleri (protonym)

References

Gmelin, J.F. 1788. Caroli a Linné systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima tertia, aucta, reformata. - pp. i–xii, 1–500. Lipsiae. (Beer). DOI: 10.5962/bhl.title.545 BHL Reference page. : 370

Vernacular names
brezhoneg: Kegin Steller
català: Gaig de Steller
čeština: Sojka Stellerova
Cymraeg: Sgrech Steller
dansk: Fyrreskade
Deutsch: Diademhäher
English: Steller's Jay
Esperanto: Stelera garolo
español: Chara de Steller
فارسی: استلرز جی
suomi: Samettinärhi
français: Geai de Steller
magyar: Bóbitás szajkó
italiano: Ghiandaia di Steller
日本語: ステラーカケス
latviešu: Stellera sīlis
Nederlands: Stellers gaai
norsk: Furuskrike
Diné bizaad: Dilzhę́ʼii
polski: Modrosójka czarnogłowa
português: Gaio-de-steller
русский: Стеллерова черноголовая голубая сойка
slovenčina: Kapuciarka diadémová
svenska: Stellerskrika
Türkçe: Steller alakargası
українська: Блакитна сойка чубата
Tiếng Việt: Giẻ cùi Steller
中文: 暗冠蓝鸦

Steller's jay (Cyanocitta stelleri) is a bird native to western North America and the mountains of Central America, closely related to the blue jay found in eastern North America. It is also known as the long-crested jay, mountain jay, and pine jay. It is the only crested jay west of the Rocky Mountains. It is also sometimes colloquially called a "blue jay" in the Pacific Northwest, but is distinct from the blue jay (C. cristata) of eastern North America. The species inhabits pine-oak and coniferous forests.

Description
Adults along the Pacific Coast have blue streaks on their black crests.

Steller's jay is about 30–34 cm (12–13 in) long and weighs about 100–140 g (3.5–4.9 oz). Steller's jay shows a great deal of regional variation throughout its range. Blackish-brown-headed birds from the north gradually become bluer-headed farther south.[3] The Steller's jay has a more slender bill and longer legs than the blue jay and, in northern populations, has a much more pronounced crest.[4]: 69 [5] It is also somewhat larger.

The head is blackish-brown, black, or dark blue, depending on the subspecies of the bird, with lighter streaks on the forehead. This dark coloring gives way from the shoulders and lower breast to silvery blue. The primaries and tail are a rich blue with darker barring. Birds in the eastern part of its range along the Great Divide have white markings on the head, especially over the eyes; birds further west have light blue markers and birds in the far west along the Pacific Coast have small, very faint, or no white or light markings at all.
Phylogeny
Cyanocitta stelleri ridgwayi, a subspecies of Steller's jay found from Chiapas to Guatemala[5]

Steller's jay (Cyanocitta stelleri) is one of two species in the genus Cyanocitta, the other species being the blue jay (Cyanocitta cristata); because the two species sometimes interbreed naturally where their ranges overlap in the eastern foothills of the Rocky Mountains, their status as distinct species has been contested. There are 18 subspecies of Steller's jays ranging from Alaska to Nicaragua, with nine found north of Mexico, often with areas of low or non-existent presence of the species separating the subspecies. At least some of the variation in the species is due to different degrees of hybridization between Steller's jays (C. stelleri) and blue jays (C. cristata).[6]

The genus Cyanocitta is part of the passerine family Corvidae, which consists of the crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, and nutcrackers.
Habitat
Western-variety Steller's jay, with all-dark head, in Alaska Maritime National Wildlife Refuge.

Steller's jay occurs in most of the forested areas of western North America as far east as the eastern foothills of the Rocky Mountains from southern Alaska in the north to northern Nicaragua in the south[6] completely replacing the blue jay prevalent on the rest of the continent in those areas. Its density is lower in the central Rocky Mountain region (Montana, Idaho, Wyoming and eastern Utah) plus the desert or scrubland areas of the Great Basin (e.g. Nevada, western Utah, southern Arizona and parts of California). Some hybridization with the blue jay in eastern foothills of the Rocky Mountains, especially Colorado, has been reported. It is also found in Mexico occurring through the interior highlands in northwestern Mexico as well as patchy populations in the rest of Mexico. In the northern end of its range it appears to be spreading from coastal Southeast Alaska across the Coast Mountains into southern Yukon Territory.[7]
Steller's jay on Pebble Beach, California, USA.

Steller's jay is also found in Mexico's interior highlands from Chihuahua and Sonora in the northwest southward to Jalisco, as well as other patchy populations found throughout Mexico. It is also found in south-central Guatemala, northern El Salvador, Honduras and Nicaragua.[3]

Although Steller's jay primarily lives in coniferous forests, it can be found in other types of forests as well. They can be found from low to moderate elevations, and on rare occasions to as high as the tree line. Steller's jays are common in residential and agricultural areas with nearby forests.[8]
Diet
Steller's jays are omnivorous and can be social with humans.

Steller's jays are omnivores; their diet is about two-thirds plant matter and one-third animal matter. They gather food both from the ground and from trees. Steller's jay's diet includes a wide range of seeds, nuts, berries and other fruit. They also eat many types of invertebrates, small rodents, eggs, and nestlings such as those of the marbled murrelet. There are some accounts of them eating small reptiles, both snakes and lizards.[8]

Acorns and conifer seeds are staples during the non-breeding season; these are often cached in the ground or in trees for later consumption. They exploit human-provided food sources, frequently scavenging picnics and campsites, where it competes with the Canada jay.

Steller's jays will visit feeders and prefer black-oil sunflower seeds, white striped sunflower seeds, cracked corn, shelled raw peanuts, and are especially attracted to whole raw peanuts. Suet is also consumed but mostly in the winter season.
Breeding

Steller's jays breed in monogamous pairs.[9] The clutch is usually incubated entirely by the female for about 16 days.[10] The male feeds the female during this time. Though they are known to be loud both day and night, during nesting they are quiet in order to not attract attention.[11]

The nest is usually in a conifer, but is sometimes built in a hollow in a tree or beneath the awning of a house or other structure. Similar in construction to the blue jay's nest, it tends to be a bit larger (25 to 43 cm (9.8 to 16.9 in)), using a number of natural materials or scavenged trash, often mixed with mud. Between two and six eggs are laid during breeding season. The eggs are oval in shape with a somewhat glossy surface. The background colour of the egg shell tends to be pale variations of greenish-blue with brown- or olive-coloured speckles.
Vocalizations

Like other jays, the Steller's jay has numerous and variable vocalizations. One common call is a harsh "SHACK-Sheck-sheck-sheck-sheck-sheck" series; another "skreeka! skreeka!" call sounds almost exactly like an old-fashioned pump handle; yet another is a soft, breathy "hoodle hoodle" whistle. Its alarm call is a harsh, nasal "wah". Some calls are sex-specific: females produce a rattling sound, while males make a high-pitched "gleep gleep".

The Steller's jay is also a noted mimic: it can imitate the vocalizations of many species of birds, other animals, and sounds of non-animal origin. It often will imitate the calls from birds of prey such as the red-tailed hawk, red-shouldered hawk, and osprey as a warning of danger to others or territorial behavior, causing other birds to seek cover and flee feeding areas.[6][8]

Steller's Jays have the ability to assess risk using different predator detection cues. They have many alarm calls that are dependent on their interaction between predator identity and cue type. Steller's Jays respond to Sharp Shinned Hawks with a longer latency to return to feeding whether they are both seen or heard. In contrast the Steller's Jays Sharp Skinned Hawks responded to Northern Goshawks with a longer latency to resume feeding if they were seen in comparison to heard. The difference in reactions suggest that this is a distinct risk of assessment being conducted by the Steller's Jays. Steller's Jays will then vary the production of their "wah", "wek", and mimic red-tailed hawk calls in response to different detection cues. [12]
Etymology

This bird is named after the German naturalist Georg Wilhelm Steller, the first European to record them, in 1741.[13][14] Johann Friedrich Gmelin formally named the species after Steller in the 13th edition of Systema Naturae (first published in 1788).
Provincial bird

The Steller's jay is the provincial bird of the Canadian province of British Columbia.[15]
References

BirdLife International (2017). "Cyanocitta stelleri". IUCN Red List of Threatened Species. 2017: e.T22705614A118809071. doi:10.2305/IUCN.UK.2017-3.RLTS.T22705614A118809071.en. Retrieved 12 November 2021.
"Cyanocitta stelleri". Integrated Taxonomic Information System. Retrieved 2019-08-13.
Walker, L. E.; Pyle, P.; Patten, M. A.; Green, E.; Davison, W.; Muehter, V. R. (2016). Rodewald, P. G. (ed.). "Steller's Jay (Cyanocitta stelleri)". The Birds of North America Online. Ithaca, New York: Cornell Lab of Ornithology. doi:10.2173/bna.343. Retrieved 2019-08-13.
Madge, S.; Burn, H. (1994). Crows and Jays: A Guide to the Crows, Jays and Magpies of the World. Boston, MA: Houghton Mifflin.
Walker, L. E.; Pyle, P.; Patten, M. A.; Greene, E.; Davison, W.; Muehter, V. R. (2020). "Steller's Jay (Cyanocitta stelleri), version 1.0". Birds of the World. doi:10.2173/bow.stejay.01. S2CID 216421604.
"Steller's Jay Cyanocitta stelleri". National Geographic. 4 May 2010. Retrieved 6 June 2013.
Ritchie, Haley (2020-10-27). "Steller's jay invasion: Coastal species making a rare appearance in southern Yukon". Yukon News. Retrieved 2020-10-27.
"Steller's Jay". Seattle Audubon Society. Retrieved 6 June 2013.
Gabriel, P. O.; Black, J. M. (July 2012). "Reproduction in Steller's Jays (Cyanocitta stelleri): individual characteristics and behavioral strategies". The Auk. 129 (3): 377–386. doi:10.1525/auk.2012.11234. JSTOR 10.1525/auk.2012.11234.
Tweit, R. C. (2005). "Steller's Jay". Texas Breeding Bird Atlas. Texas A&M. Retrieved 6 June 2013.
Kaufman, K. (13 November 2014). "Steller's Jay". Audubon Society. Retrieved 2019-08-13.
Billings, Alexis (May–June 2017). "Steller's jays assess and communicate about predator risk using detection cues and identity". Oxford Academic.
"Steller's Jay". Cornell Laboratory of Ornithology. The Birds of North America Online. Cornell University. 2013. Retrieved 6 June 2013.
Evans, Howard Ensign (1986). Halpern, Daniel (ed.). Antæus on Nature. London, UK: Collins Harvill. p. 24.

"B.C. Symbols - Province of British Columbia". gov.bc.ca. Government of British Columbia. Retrieved 2019-08-13.

Further reading
Goodwin, D. (1976). Crows of the World. Seattle, WA: University of Washington Press.
Greene, E.; Davison, W.; Davison, W.; Muehter, V.R. (1998). "Steller's jay - Cyanocitta stelleri". The Birds of North America. No. 343.

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