Superregnum: Eukaryota
Regnum: Plantae
Divisio: Tracheophyta
Divisio: Pteridophyta
Classis: Polypodiopsida
Ordo: Polypodiales
Familia: Thelypteridaceae
Subfamilia: Thelypteridoideae
Genus: Thelypteris
Species: T. confluens – T. krayanensis – T. palustris
Name
Thelypteris Schmidel, Icon. Pl. (ed. Keller): 3, 45. 1763 [18.10.1763], nom. cons. (non Thelypteris Adans. 7-8/1763 nom. rej.)
Type species: Thelypteris palustris Schott, Gen. Fil.: t. 10. 1834
Synonyms
Homotypic
Asterochlaena C.Chr., Biol. Arb. tilegn. Eng. Warming 84 (1911).
Source(s) of checklist:
Hassler, M. 2021. World Ferns. Checklist of Ferns and Lycophytes of the World. (Version 11.1 - December 2020). Thelypteris Accessed: 06 Jan 2021.
References
Hassler, M. 2020. World Ferns. Checklist of Ferns and Lycophytes of the World. (April 2020). Thelypteris Accessed: 21 Apr 2020. (Thelypteris s.str.)
Hassler, M. 2020. Thelypteris – World Ferns: Checklist of Ferns and Lycophytes of the World . In: Roskov Y., Orrell T., Nicolson D., Bailly N., Kirk P.M., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J. & Penev L. (eds.) 2020. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2020 Apr 21.
Govaerts, R. et al. 2020. Thelypteris in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2020 Apr. 21. Reference page. (Thelypteris s.l.)
Tropicos.org 2015. Thelypteris. Missouri Botanical Garden. Published online. Accessed: 16 03 2015.
Vernacular names
dansk: Mangeløv
Deutsch: Sumpffarne
English: maiden fern
suomi: Nevaimarteet
Nederlands: Moerasvaren
norsk: Myrtelgslekten
polski: Zachylnik
Thelypteris (maiden ferns[3]) is a genus of ferns in the subfamily Thelypteridoideae, family Thelypteridaceae, order Polypodiales.[1][4] Two radically different circumscriptions of the genus are in use as of January 2020. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the genus is a very small one with about two species.[1][2] In other approaches, the genus is the only one in the subfamily Thelypteridoideae, and so includes between 875[5] and 1083 species.[6]
The genus name is from Greek thēlys "female" and pteris "fern". However, "female fern" usually refers to the common lady-fern.
Taxonomy
At one time,[specify] all thelypterioid ferns were included in the genus Dryopteris because of the sorus shape. However, there are a great many differences between the groups, and these plants are now segregated in their own family.
Some researchers include the entire family Thelypteridaceae in the genus Thelypteris.[citation needed] An intermediate position is to place the bulk in Thelypteris (corresponding to Thelypterioideae of PPG I), which can then be divided into subgenera and sections corresponding to the genera of other authors, but to separate out Phegopteris and Macrothelypteris.[5] Another choice is to divide the family into many genera, one of which is Thelypteris.[2][7] This is the approach taken in the Pteridophyte Phylogeny Group classification of 2016 (PPG I).[1]
Species
Using the circumscription of the genus in PPG I,[1] as of January 2020, the Checklist of Ferns and Lycophytes of the World accepted three species:[2]
Thelypteris confluens (Thunb.) C.V.Morton
Thelypteris krayanensis K.Iwats. & M.Kato
Thelypteris palustris (Salisb.) Schott
Other sources place the entire subfamily Thelypteridoideae in the genus, and so accept many more species, including the following (synonyms from the Checklist of Ferns and Lycophytes of the World):[2]
Thelypteris aculeata A.R.Sm. = Amauropelta aculeata
Thelypteris appressa A.R.Sm. = Amauropelta appressa
Thelypteris bonapartii (Rosenst.) Alston = Amauropelta bonapartii
Thelypteris campii A.R.Sm. = Amauropelta campii
Thelypteris chimboracensis A.R.Sm. = Amauropelta chimboracensis
Thelypteris conformis (Sodiro) A.R.Sm. = Amauropelta conformis
Thelypteris correllii A.R.Sm. = Amauropelta correllii
Thelypteris dodsonii A.R.Sm. = Amauropelta dodsonii
Thelypteris elegantula (Sodiro) Alston = Amauropelta elegantula
Thelypteris euthythrix A.R.Sm. = Amauropelta euthythrix
Thelypteris fluminalis A.R.Sm. = Amauropelta fluminalis
Thelypteris inabonensis Proctor = Amauropelta inabonensis
Thelypteris kunthii (Desv.) Morton = Christella normalis
Thelypteris macra A.R.Sm. = Amauropelta macra
Thelypteris nevadensis (Baker) Clute ex Morton = Parathelypteris nevadensis
Thelypteris noveboracensis (L.) Nieuwl. = Parathelypteris noveboracensis
Thelypteris pilosa (M.Martens & Galeotti) Crawford = Stegnogramma pilosa
Thelypteris puberula (Baker) Morton = Christella puberula
Thelypteris rosenstockii (C.Chr.) R.M.Tryon = Amauropelta rosenstockii
Thelypteris semilunata (Sodiro) A.R.Sm. = Amauropelta semilunata
Thelypteris simulata (Davenport) Nieuwl. = Coryphopteris simulata
Thelypteris subtilis A.R.Sm. = Amauropelta subtilis
Thelypteris verecunda Proctor = Goniopteris verecunda
Thelypteris yaucoensis Proctor = Goniopteris yaucoensis
References
PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229. S2CID 39980610.
Hassler, Michael & Schmitt, Bernd (January 2020). "Thelypteris". Checklist of Ferns and Lycophytes of the World. 8.20. Retrieved 2020-01-11.
USDA, NRCS (n.d.). "Thelypteris". The PLANTS Database (plants.usda.gov). Greensboro, North Carolina: National Plant Data Team. Retrieved 9 December 2015.
Christenhusz, Maarten J. M.; Zhang, Xian-Chun; Schneider, Harald (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54. doi:10.11646/phytotaxa.19.1.2.
"Thelypteris". Flora of North America.
"Thelypteris Schmidel". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 2020-01-11.
Smith, Alan R.; Pryer, Kathleen M.; Schuettpelz, Eric; Korall, Petra; Schneider, Harald; Wolf, Paul G. (August 2006). "A Classification for Extant Ferns". Taxon. 55 (3): 705–31. doi:10.2307/25065646. JSTOR 25065646.
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