Isoetes

Superregnum: Eukaryota
Regnum: Plantae
Divisio: Tracheophyta
Divisio: Lycopodiophyta
Classis: Lycopodiopsida
Subclassis: Isoetidae
Ordo: Isoetales

Familia: Isoetaceae
Genus: Isoetes
Species (188)
a

I. abyssinica – I. aemulans – I. aequinoctialis – I. afloramientorum – I. alcalophila – I. alpina – I. alstonii – I. amazonica – I. anamariae – I. anatolica – I. andicola – I. andina – I. appalachiana – I. araucaniana – I. asiatica – I. asturicensis – I. attenuata – I. australis – I. azorica
b

I. baculata – I. baodongii – I. biafrana – I. bilaspurensis – I. bischlerae – I. bolanderi – I. boliviensis – I. boomii – I. boryana – I. boyacensis – I. bradei – I. brasiliensis – I. brevicula – I. butleri
c

I. cangae – I. caparaoensis – I. capensis – I. caroli – I. caroliniana – I. chubutiana – I. cipoensis – I. clavata – I. coreana – I. coromandelina – I. creussensis – I. cristata – I. cubana
d

I. delalandei – I. delilei – I. dispora – I. divyadarshanii – I. dixitii – I. drummondii – I. dubsii – I. durieui
e

I. echinospora – I. ecuadoriensis – I. ekmanii – I. elatior – I. eludens – I. engelmannii – I. escondidensis – I. eshbaughii
f

I. flaccida – I. fluitans – I. fuchsii – I. fuliginosa – I. fuscomarginata
g

I. gardneriana – I. georgiana – I. giessii – I. gigantea – I. goebelii – I. graniticola – I. gunnii – I. gymnocarpa
h

I. habbemensis – I. hallasanensis – I. hallei – I. harleyi – I. haussknechtii – I. hawaiiensis – I. heldreichii – I. hemivelata – I. herzogii – I. hewitsonii – I. hieronymi – I. histrix – I. hopei – I. howellii – I. humilior – I. hyemalis – I. hypsophila
i

I. inflata
j

I. jaegeri – I. jamaicensis – I. japonica – I. jarmaniae – I. jejuensis – I. junciformis
k

I. karstenii – I. kashyapii – I. killipii – I. kirkii
l

I. labri-draconis – I. lacustris – I. laosiensis – I. laurentiana – I. lechleri – I. libanotica – I. lithophila – I. longifolia – I. longissima – I. louisianensis – I. luetzelburgii
m

I. malinverniana – I. maritima – I. martii – I. mattaponica – I. maxima – I. melanopoda – I. melanospora – I. melanotheca – I. mexicana – I. microvela – I. minima – I. mississippiensis – I. mongerensis – I. montezumae – I. mourabaptistae – I. muelleri
n

I. naipiana – I. nana – I. narsinghgarhensis – I. neoguineensis – I. nigritiana – I. nigroreticulata – I. novogranadensis – I. nuttallii
o

I. occidentalis – I. olympica – I. orcuttii – I. organensis – I. orientalis – I. ovata
p

I. pallida – I. palmeri – I. panamensis – I. panchananii – I. panchganiensis – I. parvula – I. pedersenii – I. perrieriana – I. philippinensis – I. phrygia – I. pichisermollii – I. pirinica – I. pitotii – I. precocia – I. pringlei – I. prototypus – I. pseudojaponica – I. pusilla
q

I. quiririensis
r

I. ramboi – I. rajasthanensis – I. riparia – I. rwandensis
s

I. sabatina – I. sampathkumaranii – I. santacruzensis – I. saracochensis – I. savatieri – I. schweinfurthii – I. sehnemii – I. septentrionalis – I. serracarajensis – I. shangrilaensis – I. sinensis – I. smithii – I. spannagelii – I. spinulospora – I. stellenbossiensis – I. stephanseniae – I. stevensii – I. storkii
t

I. taiwanensis – I. tamaulipana – I. tegetiformans – I. tennesseensis – I. tenuifolia – I. texana – I. tiguliana – I. todaroana – I. toximontana – I. transvaalensis – I. triangula – I. tripus – I. tuckermanii – I. tuerckheimii
u

I. udupiensis – I. ulei
v

I. valida – I. vanensis – I. vermiculata – I. virginica – I. viridimontana
w

I. weberi – I. welwitschii – I. wormaldii
y

I. yunguiensis
Source(s) of checklist:
Unresolved

I. sahyadrii
Nothospecies

I. × angeloi – I. × altonharvillii – I. × blondeaui – I. × brittonii – I. × brochonii – I. × bruntonii – I. × carltaylori – I. × eatonii – I. × echtuckeri – I. × fairbrothersii – I. × fernaldii – I. × foveolata – I. × gopalkrishnae – I. × harveyi – I. × herb-wagneri – I. × hickeyi – I. × jeffreyi – I. × jermyi – I. × kareniae – I. × marensis – I. × michinokuana – I. × novae-angliae – I. × paratunica – I. × pseudotruncata – I. × robusta – I. × truncata
Source(s) of checklist:
List based on accepted species according to Hassler & Schmitt (2020) and Troia et al. (2016).
Fossil species

†I. bulbiformis – †I. ermayinensis – †I. reticulata
Name

Isoetes L., Sp. Pl. 2: 1100 (1753), Gen. Pl. ed. 5: 486 (1754) (optional spelling: Isoëtes, see Art. 60.6 ICN).

Lectotype species: Isoetes lacustris L. (designated by Jonsell & Jarvis, Reg. Veg. 127: 57 (1993).)

Synonyms

Heterotypic
Calamaria Boehm., Def. Gen. Pl. ed. Boehmer 500 (1760). nom. illeg.
Cephaloceraton Gennari, Comment. Soc. Critt. Ital. 1: 111 (1862).
Type species: Cephaloceraton histrix (Bory & Durieu) Gennari
Isoetella Gennari, Comment. Soc. Critt. Ital. 1: 114 (1862).
Type species: Isoetella durieui (Bory) Gennari
Sivetes Raf., Med. Repos. 11. 5: 358 (1808).
Stylites Amstutz, Ann. Missouri Bot. Gard. 44: 121 (1957).
Typus: Stylites andicola Amstutz, Ann. Missouri Bot. Gard. 44: 121 (1957).
Subularia Ray, Syn. Meth. Stirp. Brit.: 283 (1696). nom. inval. (Pre-Linnaean)

Distribution
Native distribution areas:

Continental: Europe
Regional: Northern Europe
Denmark, Finland, Great Britain, Iceland, Ireland, Norway, Sweden.
Regional: Middle Europe
Austria, Belgium, Czechoslovakia, Germany, Netherlands, Poland, Switzerland.
Regional: Southwestern Europe
Baleares, Corse, France, Portugal, Sardegna, Spain.
Regional: Southeastern Europe
Bulgaria, Greece, Italy, Kriti, Sicilia, Turkey-in-Europe, Yugoslavia.
Regional: Eastern Europe
Baltic States, Central European Russia, East European Russia, North European Russia.
Continental: Africa
Regional: Northern Africa
Algeria, Libya, Morocco, Tunisia.
Regional: Macaronesia
Azores.
Regional: West Tropical Africa
Benin, Burkina, Ghana, Guinea, Ivory Coast, Mali, Nigeria, Niger, Senegal.
Regional: West-Central Tropical Africa
Burundi, Central African Republic, Cameroon, Gulf of Guinea Islands.
Regional: Northeast Tropical Africa
Chad, Eritrea, Ethiopia, Sudan.
Regional: East Tropical Africa
Kenya, Tanzania.
Regional: South Tropical Africa
Angola, Malawi, Mozambique, Zambia, Zimbabwe.
Regional: Southern Africa
Botswana, Cape Provinces, Lesotho, Namibia, KwaZulu-Natal, Free State, Swaziland, Northern Provinces.
Regional: Western Indian Ocean
Madagascar.
Continental: Asia-Temperate
Regional: Siberia
Altay, Buryatiya, Chita, Irkutsk, West Siberia.
Regional: Russian Far East
Kamchatka, Kuril Islands, Primorye, Sakhalin.
Regional: Western Asia
East Aegean Islands, Lebanon-Syria, Turkey.
Regional: China
China South-Central, China Southeast.
Regional: Eastern Asia
Japan, Korea, Taiwan.
Continental: Asia-Tropical
Regional: Indian Subcontinent
India, Nepal, Sri Lanka, West Himalaya.
Regional: Indo-China
Laos, Myanmar, Thailand, Vietnam.
Regional: Malesia
Philippines, Sulawesi.
Regional: Papuasia
New Guinea.
Continental: Australasia
Regional: Australia
New South Wales, Northern Territory, Queensland, South Australia, Tasmania, Victoria, Western Australia.
Regional: New Zealand
New Zealand North, New Zealand South.
Continental: Pacific
Regional: Northwestern Pacific
Caroline Islands.
Regional: North-Central Pacific
Hawaii.
Continental: Northern America
Regional: Subarctic America
Aleutian Islands, Alaska, Greenland, Nunavut, Northwest Territories, Yukon.
Regional: Western Canada
Alberta, British Columbia, Manitoba, Saskatchewan.
Regional: Eastern Canada
Labrador, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Island, Québec.
Regional: Northwestern U.S.A.
Colorado, Idaho, Montana, Oregon, Washington, Wyoming.
Regional: North-Central U.S.A.
Illinois, Iowa, Kansas, Minnesota, Missouri, Nebraska, Oklahoma, South Dakota, Wisconsin.
Regional: Northeastern U.S.A.
Connecticut, Indiana, Maine, Massachusetts, Michigan, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Vermont, West Virginia.
Regional: Southwestern U.S.A.
Arizona, California, Nevada, Utah.
Regional: South-Central U.S.A.
New Mexico, Texas.
Regional: Southeastern U.S.A.
Alabama, Arkansas, Delaware, Florida, Georgia, Kentucky, Louisiana, Maryland, Mississippi, North Carolina, South Carolina, Tennessee, Virginia, District of Columbia.
Regional: Mexico
Mexico Central, Mexico Northeast, Mexico Northwest, Mexico Southwest, Mexico Southeast.
Continental: Southern America
Regional: Central America
Belize, Costa Rica, Guatemala, Nicaragua.
Regional: Caribbean
Cuba, Dominican Republic, Haiti, Jamaica.
Regional: Northern South America
French Guiana, Guyana, Venezuela.
Regional: Western South America
Bolivia, Colombia, Ecuador, Peru.
Regional: Brazil
Brazil West-Central, Brazil Northeast, Brazil Southeast, Brazil North, Brazil South.
Regional: Southern South America
Argentina Northeast, Argentina South, Argentina Northwest, Chile Central, Chile South, Paraguay, Uruguay.

References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition
References
Primary references

Linnaeus, C. 1753. Species Plantarum. Tomus II: 1100. Reference page.
Linnaeus, C. 1754. Genera Plantarum, ed. 5: 486. Reference page.

Additional references

Pfeiffer, N.E. 1922. Monograph of the Isoetaceae. Annals of the Missouri Botanical Garden 9(2): 79–218, pls. xii–xix. JSTOR, BHL Reference page.
Pteridophyte Phylogeny Group (PPG I) 2016. A community-derived classification for extant lycophytes and ferns. Journal of Systematics and Evolution 54(6): 563–603. DOI: 10.1111/jse.12229 Reference page.
Troìa, A., Pereira, J.B.S., Kim, C. & Taylor, W.C. 2016. The genus Isoetes (Isoetaceae): a provisional checklist of the accepted and unresolved taxa. Phytotaxa 277(2): 101–145. DOI: 10.11646/phytotaxa.277.2.1 ResearchGate Reference page.
Brunton, D.F. & Troìa, A. 2018. Global Review of recent taxonomic research into Isoetes (Isoetaceae), with implications for biogeography and conservation. Fern Gazette 20(8): 309–333. ResearchGate Reference page.
Troìa, A. & Rouhan, G.. 2018. Clarifying the nomenclature of some Euro-Mediterranean quillworts (Isoetes, Isoetaceae): Indicator species and species of conservation concern. Taxon 67(5): 996–1004. DOI: 10.12705/675.10 Reference page.
Singh, S.K., Shukla, P.K., Brunton, D.F., Dubey, N.K. & Shukla, S.K. 2020. The taxonomy and conservation status of Isoetes (Isoetaceae; Lycopodiopsida) in India. Botany Letters 168(2): 200–226. DOI: 10.1080/23818107.2020.1848621. Reference page.

Links

Swale, B. & Hassler, M. 2002. Checklist of World Ferns. Family Isoetaceae, genus Isoetes, world species list.
Hassler, M. 2017. World Ferns. Checklist of Ferns and Lycophytes of the World. (Version 7.20 – October 2017). Isoetes Accessed: 20 Oct 2017.
International Plant Names Index. 2017. Isoetes. Published online. Accessed: 6 Nov 2017.
Tropicos.org 2017. Isoetes. Missouri Botanical Garden. Published online. Accessed: 18 8 2017.

Vernacular names
беларуская: Палушнік
dansk: rasenføde
Deutsch: Brachsenkräuter
English: Quillwort
eesti: Lahnarohi
suomi: Lahnanruohot
français: Isoète
magyar: Durdafű
македонски: Шилатични
Nederlands: Biesvaren
norsk: Brasmegrasslekten
polski: Poryblin
svenska: Braxengräs
中文: 水韭