Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Caenogastropoda
Ordo: Littorinimorpha
Superfamilia: Truncatelloidea
Familia: Hydrobiidae
Subfamiliae (11): Belgrandiellinae – Belgrandiinae – Caspiinae – Horatiinae – Hydrobiinae – Islamiinae – Mercuriinae – Nymphophilinae – Pseudamnicolinae – Pyrgulinae – Sadlerianinae – Shadiniinae
Genera Incertae sedis (20 + 62†): Achaiohydrobia – Anatolidamnicola – Antillobia – Brasovia – Bullaregia – Hemite – Heterocyclus – Isparta – Lartetia – Navalis – Parhydrobia – Probythinella – Pseudopaludinella – Salaeniella – Sivasi – Stygobium – Syrofontana – Tefennia – Torosia – Travunijana – †Acrostele – †Arabiella – †Baglivia – †Bania – †Banneina – †Beogradica – †Briardia – †Charydrobia – †Cirsomphalus – †Coelacanthia – †Costhyrella – †Ctyrokya – †Goniatogyra – †Goniochilus – †Gyromelania – †Illyricella – †Iraklimelania – †Jekeliella – †Kadolskya – †Kerchia – †Lapparentia – †Lavansia – †Liaoheniella – †Lisinskia – †Lutetiella – †Lysiogyrus – †Microbeliscus – †Micromelania – †Micropyrgus – †Mikrogoniochilus – †Mirolaminatus – †Montjavoultia – †Muellerpalia – †Nannopyrgula – †Nematurella – †Neohoratia – †Odontohydrobia – †Parateinostoma – †Pechelbronnia – †Pontohydrobia – †Potamaclis – †Potamolithoides – †Probaicalia – †Prososthenia – †Pyrgulella – †Renistoma – †Rhodopyrgula – †Robicia – †Saccoia – †Salakosia – †Salalahia – †Scalimelania – †Sellia – †Socenia – †Staja – †Stalioa – †Staliopsis – †Tournouerina – †Unicarinata – †Vrazia – †Wallaua – †Xestopyrguloides
Overview of genera (228)
A
Achaiohydrobia – Adriohydrobia – Agrafia – Alzoniella – Anagastina – Anatolidamnicola – Andrusovia – Antibaria – Antillobia – Arganiella – Avenionia – †Acrostele – †Arabiella
B
Balkanica – Balkanospeum – Belgrandia – Belgrandiella – Birgella – Boetersiella – Bracenica – Brasovia – Bucharamnicola – Bullaregia – †Baglivia – †Bania – †Banneina – †Belchatovia – †Beogradica – †Briardia
C
Caspia – Cavernisa – Chilopyrgula – Chirgisia – Chondrobasis – Cilgia – Cincinnatia – Clathrocaspia – Clessiniola – Corbellaria – Corrosella –}} †Charydrobia – †Cirsomphalus – †Coelacanthia – †Costhyrella – †Ctyrokya – †Cyclothyrella
D
Dalmatella – Daphniola – Deganta – Devetakia – Devetakiola – Dianella – Diegus
E
Ecrobia
F
Falniowskia – Falsipyrgula – Fissuria – Floridobia
G
Ginaia – Globuliana – Gloeria – Gocea – Graecoanatolica – Graecoarganiella – Graecorientalia – Graziana – Grossuana – Guadiella – †Goniatogyra – †Goniochilus – †Graecamnicola – †Gyromelania
H
Hadziella – Hauffenia – Hemite – Heraultiella – Heterocyclus – Horatia – Hoyia – Hydrobia
I
Iberhoratia – Insignia – Intermaria – Isimerope – Islamia – Isparta – Istriana – Iverakia – †Illyricella – †Iraklimelania
J
Josefus – †Jekeliella
K
Karucia – Kaskakia – Kerkia – Kolevia – †Kadolskya – †Kerchia
L
Laevicaspia – Lanzaiopsis – Lartetia – Litthabitella – Lyhnidia – †Lapparentia – †Lavansia – †Liaoheniella – †Lisinskia – †Lutetiella – †Lysiogyrus
M
Malaprespia – Marstonia – Martensamnicola – Mercuria – Meyrargueria – Micropyrgula – Microstygia – Milesiana – Motsametia – Myrtoessa – †Maeotidia – †Marticia – †Martinietta – †Microbeliscus – †Micromelania – †Microprososthenia – †Micropyrgus – †Mikrogoniochilus – †Mirolaminatus – †Montjavoultia – †Muellerpalia
N
Narentiana – Navalis – Neofossarulus – Nicolaia – Notogillia – †Nannopyrgula – †Nematurella – †Neohoratia
O
Ohridohauffenia – Ohridohoratia – Ohridopyrgula – Ohrigocea – †Odontohydrobia
P
Palacanthilhiopsis – Parhydrobia – Pauluccinella – Peringia – Persipyrgula – Pezzolia – Phreatica – Plagigeyeria – Plesiella – Polinskiola – Pontobelgrandiella – Pontohoratia – Prespiana – Prespolitorea – Probythinella – Pseudamnicola – Pseudavenionia – Pseudohoratia – Pseudoislamia – Pseudopaludinella – Pyrgohydrobia – Pyrgula – Pyrgulopsis – †Parateinostoma – †Pechelbronnia – †Pontohydrobia – †Potamaclis – †Potamolithoides – †Probaicalia – †Prososthenia – †Pseudodianella – †Pyrgulella
R
Radomaniola – Rhapinema – †Renistoma – †Rhodopyrgula – †Robicia – †Romania
S
Sadleriana – Salaeniella – Salenthydrobia – Sarajana – Sardohoratia – Sarkhia – Saxurinator – Shadinia – Sivasi – Sogdamnicola – Spathogyna – Spilochlamys – Stankovicia – Stiobia – Stoyanovia – Strugia – Stygobium – Sumia – Syrofontana – †Saccoia – †Salakosia – †Salalahia – †Scalimelania – †Sellia – †Sibiropyrgula – †Socenia – †Staja – †Stalioa – †Staliopsis
T
Tadzhikamnicola – Tarraconia – Tefennia – Terranigra – Torosia – Trachyochridia – Travunijana – Trichonia – Turcorientalia – Turkmenamnicola – Turricaspia – †Tournoueria – †Tournouerina
U
Ulskia – †Unicarinata
V
Valvatamnicola – Vinodolia – †Vrazia
W
†Wallaua
X
Xestopyrgula – †Xestopyrguloides
Z
Zaumia – Zeteana
Name
Hydrobiidae Stimpson, 1865
References
Template:Stimpson, 1865
Becker, M., Zielske, S. & Haase, M. 2016. Conflict of mitochondrial phylogeny and morphology-based classification in a pair of freshwater gastropods (Caenogastropda, Truncatelloidea, Tateidae) from New Caledonia. ZooKeys 603: 17–32. DOI: 10.3897/zookeys.603.9144. Reference page.
Boulaassafer, K., Ghamizi, M. & Delicado, D. 2018. The genus Mercuria Boeters, 1971 in Morocco: first molecular phylogeny of the genus and description of two new species (Caenogastropoda, Truncatelloidea, Hydrobiidae). ZooKeys 782: 95–128. DOI: 10.3897/zookeys.782.26797 Reference page.
Falniowski, A., Grego, J., Rysiewska, A., Osikowski, A. & Hofman, S. 2021. A new genus and species of Hydrobiidae Stimpson, 1865 (Caenogastropoda, Truncatelloidea) from Peloponnese, Greece. Zookeys 1037: 161–179. DOI: 10.3897/zookeys.1037.64038 Open access Reference page.
Falniowski, A. & Sarbu, S.M. 2015. Two new Truncatelloidea species from Melissotrypa Cave in Greece (Caenogastropoda). Zookeys 530: 1–14. DOI: 10.3897/zookeys.530.6137 Full article Reference page.
Glöer, P. & Pešić, V. 2014: New subterranean freshwater gastropods of Montenegro (Mollusca: Gastropoda: Hydrobiidae). Ecologica Montenegrina 1(2): 82-88. full article (PDF). Reference page.
Kabat, A.R.; Hershler, R. 1993: The Prosobranch Snail Family Hydrobiidae (Gastropoda: Rissooidea): Review of Classification and Supraspecific Taxa. Smithsonian contributions to zoology, (547)
Hershler, R.; Liu, H.-P. 2010: Two new, possibly threatened species of Pyrgulopsis (Gastropoda: Hydrobiidae) from southwestern California. Zootaxa, 2343: 1–17. Preview
Hershler, R., Liu, H-P. & Hubbart, N. 2017. Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F. multifarius. ZooKeys 679: 1—20. DOI: 10.3897/zookeys.679.13472. Reference page.
Liu, H-P. & Hershler, R. 2019. A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California–Oregon. Zookeys, 812: 47–67. DOI: 10.3897/zookeys.812.29205 Reference page.
Liu, H-P., Hershler, R. & Rossel, C.S. 2015. Taxonomic status of the Columbia duskysnail (Truncatelloidea, Amnicolidae, Colligyrus). Zookeys 514: 1–13. DOI: 10.3897/zookeys.514.9919 Full article Reference page.
Neubauer, T.A., van de Velde, S., Yanina, T. & Wesselingh, F.P. 2018. A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys 770: 43–103. DOI: 10.3897/zookeys.770.25365 Reference page.
Radea, C., Parmakelis, A. & Giokas, S. 2016. Myrtoessa hyas, a new valvatiform genus and a new species of the Hydrobiidae (Caenogastropoda, Truncatelloidea) from Greece. ZooKeys 640: 1-18. DOI: 10.3897/zookeys.640.10674. Reference page.
Links
Hydrobiidae in the World Register of Marine Species
https://web.archive.org/ Catalogue of Life: 2019 Annual Checklist
Vernacular names
English: mud snails
Hydrobiidae, commonly known as mud snails, is a large cosmopolitan family of very small freshwater and brackish water snails with an operculum; they are in the order Littorinimorpha. [1]
Distribution
Hydrobiidae are found in much of the world, inhabiting all continents except Antarctica. In Australia alone there are over 260 species in the family.[2]
Description
These are very small or minute snails, with a shell height of less than 8 mm. The dextrally-coiled shells are smooth (except for growth lines conforming to the shape of the outer lip) and are usually rather nondescript. The shell offers very few robust characteristics to the systematist who is attempting to classify the species within this family. This difficulty is compounded by a high degree of intraspecific variation. Descriptions often have to be based on the characteristics of the operculum, radula and penis.
The shell of species within this family varies from planispiral to needle-shaped. The shell may have an open umbilicus or a plugged umbilicus. The thickness of the shell can vary from thin to fairly solid. The shell may be transparent and horn-colored, or colorless.
The number of whorls in the shell varies between two and eight. The shell can sometimes even assume a corkscrew or hornlike shape by loosening of the attachment of body whorl. The periostracum (outer layer of the shell) is usually thin, and is often colored. It can sometimes show hair-like projections.
The sinuous aperture is entire, not interrupted by a siphonal canal or other extensions. It is sometimes thickened. The protoconch is usually shaped like a dome, and usually shows a few spirals.
The corneous operculum is usually well-formed and shows only a few spirals.
The head, foot, mantle and visceral coil are colored pale gray to dark purple-black with melanin pigments. Subterranean species are often unpigmented.
The strong foot can be retracted into the shell. The mucous glands are situated at the anterior edge of the foot. There are no posterior mucous glands. The symmetrical cephalic tentacles are threadlike, with blunt or rounded tips. The eyes are located at the base of the tentacles.
The ctenidium (a comblike respiratory apparatus) goes along most of the length of the pallial cavity. This respiratory gill consists of 10 to 200 triangular filaments. The osphradium, the olfactory organ linked with the respiration organ, is usually relatively small.
The taenioglossate radula consists of more than fifty rows of teeth. The central tooth is trapezoidal. The lateral teeth have few to numerous cusps. The marginal teeth are usually with numerous cusps.
The species usually have both male and female individuals, but very rarely reproduction may be parthenogenic, caused by internal fertilization. The females lay eggs in single capsules on the leaves or stems of water plants. But sometimes they produce eggs that are hatched within the pallial gonoduct of the body, and in these cases the young are born alive. Species that inhabit estuaries sometimes produce veliger larvae.
The fossil record of this family extends back to the Early Carboniferous.[3]
Ecology
Habitat
Most species of this family live in freshwater (lakes, ponds, rivers, streams), but some are found in brackish water or at the borders between freshwater and brackish water. A few occur in marine environments on sandy or muddy bottoms between algae and sea grass.
Feeding habits
These small snails feed on algae, diatoms and detritus.
Taxonomy
This is the largest family within the superfamily Truncatelloidea. At one time or another some 400 genera have been assigned to this family, and probably more than 1,000 species.[3] The Hydrobiidae family is the most widespread and diverse family of freshwater molluscs in the world, occupying a range of habitats from streams and estuarine creeks to alpine bogs.
This family was originally named by Troschel in 1857, as the group Hydrobiae. Troschel was not certain of their rank, and he placed them in the Taenioglossata: Ctenobranchiata between the Lithoglyphi and Ancyloti.[3] Over the years there have been numerous attempts to give an adequate and more finely divided classification. Thiele (1925, 1929 and 1931) set up the most comprehensive classification, with a review of the family at generic level.
Notable works about taxonomy of Hydrobiidae includes works by Radoman, for example Radoman (1983).[4]
The classification, as specified by Kabat and Hershler (1993),[3] does not form a monophyletic group, and was in need of revision.
A study by Wilke et al. (2001)[5] using molecular data from COI (Cytochrome c oxidase subunit I) and 18S genes has resulted in a new tentative set of subfamilies: Hydrobiinae, Pseudamnicolinae, Nymphophilinae, Islamiinae and Horatiinae.
2005 taxonomy
The following subfamilies are recognized in the gastropod taxonomy of Bouchet & Rocroi from 2005:[6]
Hydrobiinae Stimpson, 1865 – synonyms: Paludestrinidae Newton, 1891; Pyrgorientaliinae Radoman, 1977; Pseudocaspiidae Sitnikova & Starobogatov, 1983
Belgrandiinae de Stefani, 1877 – synonyms: Horatiini D. W. Taylor, 1966; Sadlerianinae Radoman, 1973; Pseudohoratiinae Radoman, 1973; Orientaliidae Radoman, 1973 (inv.); Lithoglyphulidae Radoman, 1973; Orientalinidae Radoman, 1978 (inv.); Belgrandiellinae Radoman, 1983; Dabrianidae Starobogatov, 1983; Istrianidae Starobogatov, 1983; Kireliinae Starobogatov, 1983; lanzaiidae Starobogatov, 1983; Tanousiidae Starobogatov, 1983; Bucharamnicolinae Izzatulaev, Sitnikova & Starobogatov, 1985; Martensamnicolinae Izzatulaev, Sitnikova & Starobogatov, 1985; Turkmenamnicolinae Izzatulaev, Sitnikova & Starobogatov, 1985
Clenchiellinae D. W. Taylor, 1966[7]
Islamiinae Radoman, 1973
Navarriellinae García-Guerrero, Miller & Ramos, 2023
Nymphophilinae D. W. Taylor, 1966[7]
Pseudamnicolinae Radoman, 1977
Pyrgulinae Brusina, 1882 (1869)[8] – synonyms: Caspiidae B. Dybowski, 1913; Microliopalaeinae B. Dybowski & Grochmalicki, 1914; Micromelaniidae B. Dybowski & Grochmalicki, 1914; Turricaspiinae B. Dybowski & Grochmalicki, 1915; Liosarmatinae B. Dybowski & Grochmalicki, 1920; Chilopyrgulinae Radoman, 1973; Micropyrgulidae Radoman, 1973; Falsipyrgulinae Radoman, 1983; Ohridopyrgulinae Radoman, 1983; Prosostheniinae Pana, 1989
Tateinae Thiele, 1925[9] – synonyms: Potamopyrgidae F. C. Baker, 1928; Hemistomiinae Thiele, 1929
The Amnicolidae and Cochliopidae are considered as distinct families according to the taxonomy of Bouchet & Rocroi (2005).[6]
Genera
Belgrandiellinae Radoman, 1983
Aetis Chertoprud, Grego & Mumladze, 2023
Andrusovia Brusina, 1903
Arganiella Giusti & Pezzoli, 1980
Balkanica Georgiev, 2011
Balkanospeum Georgiev, 2012
Belgrandiella A. J. Wagner, 1928
Boleana Radoman, 1975: synonym of Belgrandiella A. J. Wagner, 1928
Bucharamnicola Izzatullaev, Sitnikova & Starobogatov, 1985
Cavernisa Radoman, 1978
Chirgisia Glöer, Boeters & Pešić, 2014
Cilgia Schütt, 1968
Colchiella Chertoprud, Grego & Mumladze, 2023
Costellina Kuščer, 1933: synonym of Paladilhiopsis Pavlović, 1913
Graziana Radoman, 1975
Heraultiella Bodon, Manganelli & Giusti, 2002
Insignia Angelov, 1972
Istriana Velkovrh, 1971
Iverakia Glöer & Pešić, 2014
Kerkia Radoman, 1978
Kolevia Georgiev & Glöer, 2015
Meyrargueria Girardi, 2009
Microstygia Georgiev & Glöer, 2015
Motsametia Vinarski, Palatov & Glöer, 2014
Palacanthilhiopsis Bernasconi, 1988
Phreatica Velkovrh, 1970
Pontohoratia Vinarski, Palatov & Glöer, 2014
Sarajana Radoman, 1975
Sataplia Chertoprud, Grego & Mumladze, 2023
Sogdamnicola Izzatullaev, Sitnikova & Starobogatov, 1984
Tadzhikamnicola Izzatullaev, 2004
Tschernomorica Vinarski & Palatov, 2019
Turkmenamnicola Izzatullaev, Sitnikova & Starobogatov, 1985
Valvatamnicola Izzatullaev, Sitnikova & Starobogatov, 1984
Zeteana Glöer & Pešić, 2014
Belgrandiinae de Stefani, 1877
Antibaria Radoman, 1983
†Belchatovia Kadolsky & Piechocki, 2000
Belgrandia Bourguignat, 1870
Bracenica Radoman, 1973
†Cyclothyrella Neubauer, Mandic, Harzhauser & Hrvatović, 2013
Dalmatella Velkovrh, 1970
Daphniola Radoman, 1973
Gloeria Georgiev, Dedov & Cheshmedjiev, 2012
Gocea Hadžišče, 1956
Graecorientalia Radoman, 1983
Grossuana Radoman, 1983
Guadiella Boeters, 2003
Hadziella Kuščer, 1932
Isimerope Radea & Parmakelis, 2013
Karucia Glöer & Pešić, 2013
Litthabitella Boeters, 1970
Lyhnidia Radoman, 1962
Malaprespia Radoman, 1973
Martensamnicola Izzatullaev, Sitnikova & Starobogatov, 1985
†Martinietta Schlickum, 1974
†Microprososthenia Kadolsky & Piechocki, 2000
Myrtoessa Radea, 2016
Narentiana Radoman, 1973
Ohridohauffenia Hadžišče, 1959
Ohridohoratia Hadžišče, 1959
Ohrigocea Hadžišče, 1959
Pauluccinella Fo. Giusti & Pezzoli, 1990
Plesiella Boeters, 2003
Prespiana Radoman, 1973
Prespolitorea Radoman, 1983
Pseudavenionia Bodon & Giusti, 1982
Pseudohoratia Radoman, 1967
Pseudoislamia Radoman, 1979
Strugia Radoman, 1973
Sumia Glöer & Mrkvicka, 2015
Tarraconia Ramos & Arconada, 2000
Trichonia Schütt, 1980
Turcorientalia Radoman, 1973
Zaumia Radoman, 1983
Zavalia Radoman, 1974
Bullaregiinae Delicado, Hauffe & Wilke, 2023 (unavailable name, description in Supplementary Material does not fulfill the requirements of the Code for e-only publications )
Belgrandiellopsis Khalloufi, Béjaoui & Delicado, 2020
Biserta Khalloufi, Béjaoui & Delicado, 2020
Bullaregia Khalloufi, Béjaoui & Delicado, 2017
Caspiinae B. Dybowski, 1913
† Alizadella Kabakova, 1966
† Azeria Kabakova, 1967
Caspia Clessin & W. Dybowski, 1887
Clathrocaspia Lindholm, 1930
Erosiconcha Delicado & Gürlek, 2021
Graecoanatolica Radoman, 1973
Ulskia Logvinenko & Starobogatov, 1969
Horatiinae D.W. Taylor, 1966
Anagastina Radoman, 1978
Caucasogeyeria Grego & Mumladze, 2020
Caucasopsis Grego & Mumladze, 2020
Dalmatinella Radoman, 1973
Daudebardiella O. Boettger, 1905
Graecoarganiella Falniowski & Szarowska, 2011
Hausdorfenia Grego & Mumladze, 2020
Horatia Bourguignat, 1887
Iglicopsis Falniowski & Hofman, 2021
Imeretiopsis Grego & Mumladze, 2020
Kartvelobia Grego & Mumladze, 2020
Lerniana Delicado & Hauffe, 2022
Montenegrospeum Pešić & Glöer, 2013
Pezzolia Bodon & Giusti, 1986
Radomaniola Szarowska, 2007
Sadleriana Clessin, 1890
Sardohoratia Manganelli, Bodon, Cianfanelli, Talenti & Giusti, 1998
Tanousia Servain, 1881
Travunijana Grego & Glöer, 2019
Vinodolia Radoman, 1973
A shell of Peringia ulvae
Hydrobiinae Stimpson, 1865
Achaiohydrobia Falniowski, 2021
Adriohydrobia Radoman, 1977
Cariohydrobia Delicado & Gürlek, 2021
Ecrobia Stimpson, 1865
† Harzhauseria Neubauer & Wesselingh, 2023
Hoyia F. C. Baker, 1926
Hydrobia W. Hartmann, 1821
Peringia Paladilhe, 1874
† Polycirsus Cossmann, 1888
†Romania Cossmann, 1913
Salenthydrobia Wilke, 2003
† Tournoueria Brusina, 1870
Islamiinae Radoman, 1973
Actenidia Arconada & Ramos, 2024
Alzoniella Giusti & Bodon, 1984
Aretiana Delicado & Ramos, 2021
Avenionia Nicolas, 1882
Beatrix J. P. Miller, García-Guerrero, Delicado & Ramos, 2024
Boetersiella Arconada & Ramos, 2001
Chondrobasis Arconada & Ramos, 2001
Corbellaria Callot-Girardi & Boeters, 2012
Deganta Arconada & Ramos, 2019
Fissuria Boeters, 1981
Hauffenia Pollonera, 1898
Iberhoratia Arconada & Ramos, 2007
Islamia Radoman, 1973
Josefus Arconada & Ramos, 2006
Milesiana Arconada & Ramos, 2006
Spathogyna Arconada & Ramos, 2002
Mercuriinae Boeters & Falkner, 2017
Mercuria Boeters, 1971
Navarriellinae García-Guerrero, Miller & Ramos, 2023
Navarriella Boeters, 2000
Nymphophilinae
Birgella Baker, 1926
Cincinnatia Pilsbry, 1891
Floridobia F. G. Thompson & Hershler, 2002
Marstonia F. C. Baker, 1926
Notogillia Pilsbry, 1953
Pyrgulopsis Call & Pilsbry, 1886
Rhapinema F. G. Thompson, 1970
Spilochlamys F. G. Thompson, 1968
Stiobia F. G. Thompson, 1978
Pontobelgrandiellinae Delicado, Hauffe & Wilke, 2023 (unavailable name, description in Supplementary Material does not fulfill the requirements of the Code for e-only publications)
Cavernisa Radoman, 1978
Devetakia Georgiev & Glöer, 2011
Devetakiola Georgiev, 2017
Pontobelgrandiella Radoman, 1978
Stoyanovia Georgiev, 2017
Pseudamnicolinae Radoman, 1977
Corrosella Boeters, 1970
Diegus Delicado, Machordom & Ramos, 2016
Falniowskia Bernasconi, 1991
†Graecamnicola Willmann, 1981
Kaskakia Glöer & Pešić, 2012
Kitrinomatousa Glöer & Kolokotronis, 2023
Pseudamnicola Paulucci, 1878
Sarkhia Glöer & Pešić, 2012
Pyrgulinae E. von Martens, 1858
† Aenigmapyrgus Popova & Starobogatov, 1970
† Annulifer Cossmann, 1921
Chilopyrgula Brusina, 1896
Clessiniola Lindholm, 1924
Dianella Gude, 1913
Falsipyrgula Radoman, 1973
Ginaia Brusina, 1896
† Itomelania K. Suzuki, 1943
Laevicaspia Dybowski & Grochmalicki, 1917
†Maeotidia Andrusov, 1890
†Marticia Brusina, 1897
† Micromelania Brusina, 1874
Micropyrgula Polinski, 1929
Neofossarulus Poliński, 1929
Ohridopyrgula Radoman, 1983
†Pseudodianella Neubauer, Mandic, Harzhauser & Hrvatović, 2013
Pyrgohydrobia Radoman, 1955
Pyrgorientalia Radoman, 1973
Pyrgula de Cristofori & Jan, 1832
† Sibiropyrgula Lindholm, 1932
Stankovicia Poliński, 1929
Trachyochridia Poliński, 1929
Turricaspia B. Dybowski & Grochmalicki, 1915
Xestopyrgula Poliński, 1929
Shadiniinae V. Anistratenko, Peretolchina, Sitnikova & Palatov, 2017
Intermaria Delicado, Pešić & Glöer, 2016
Nicolaia Glöer, Bößneck, Walther & Neiber, 2016
Persipyrgula Delicado, Pešić & Glöer, 2016
Shadinia Akramowski, 1976
Incertae sedis
Agrafia Szarowska & Falniowski, 2011
Anatolidamnicola (sic): synonym of Anadoludamnicola Şahin, Koca & Yıldırım, 2012 (incorrect original spelling)
Antillobia Altaba, 1993: synonym of Pyrgophorus Ancey, 1888 (subjective junior synonym)
†Arabiella Kadolsky, Harzhauser & Neubauer in Harzhauser et al., 2016
Austropyrgus Cotton, 1942
†Baglivia Brusina, 1892
†Bania Brusina, 1896
†Beogradica Pavlović, 1927
Berkanella Taybi, Glöer & Mabrouki, 2024
† Bernicia L. R. Cox, 1927
†Brasovia Neubauer, Kroh, Harzhauser, Georgopoulou & Mandic, 2015
† Briardia Munier-Chalmas, 1884
Bullaregia Khalloufi, Béjaoui & Delicado, 2017
Caspiohydrobia Starobogatov, 1970 (nomen dubium)
† Charydrobia Stache, 1889
† Chazarella A. A. Ali-Zade, 1967
† Cirsomphalus Cossmann, 1907
†Coelacanthia Andrusov, 1890
† Costhyrella Pacaud, 1997
†Ctyrokya Schlickum, 1965
† Edrozeba Guzhov, 2022
Falsibelgrandiella Radoman, 1973
Fessia Glöer, Mabrouki & Taybi, 2020
† Fuxinia X.-H. Yu, 1982
Gafaita Taybi, Glöer & Mabrouki, 2023
† Goniatogyra Cossmann, 1921
†Goniochilus Sandberger, 1875
†Gyromelania Wenz, 1939
Heideella Backhuys & Boeters, 1974
Hemite Gürlek, 2018
Heterocyclus Crosse, 1872
Idrisiella Mabrouki, Glöer & Taybi, 2022
Ifrania Glöer, Mabrouki & Taybi, 2020
†Illyricella Neubauer, Mandic & Harzhauser, 2016
†Iraklimelania Willmann, 1981
† Islamiella Guzhov, 2023
†Jekeliella Bandel, 2010
†Kadolskya Neubauer & Harzhauser, 2016
Kaskakia Glöer & Pešić, 2012
†Kerchia Bandel, 2010
†Lisinskia Brusina, 1897
Littoridinops Pilsbry, 1952
†Lutetiella Kadolsky, 2015
† Lyobasis L.-S. Huang, 1987 (accepted > unreplaced junior homonym, junior homonym of Lyobasis Pilsbry, 1903)
Mahrazia Mabrouki, Taybi & Gloër, 2022
Menzella Mabrouki, Glöer & Taybi, 2023
†Microbeliscus Sandberger, 1875
†Micromelania Brusina, 1874
†Mikrogoniochilus Willmann, 1981
† Mirolaminatus H.-J. Wang, 1977
† Montjavoultia Raspail, 1909
Motsametia Vinarski, Palatov & Glöer, 2014
†Muellerpalia Bandel, 2010
Navalis Quiñonero-Salgado & Rolán, 2017
†Nematurella Sandberger, 1875
Neohoratia Schütt, 1961
†Odontohydrobia Pavlović, 1927
Ohridonaumia Glöer & Pešić, 2023
† Parateinostoma Oppenheim, 1892
Parhydrobia Cossmann & Dollfus, 1913
† Pechelbronnia R. Janssen, 2007
Phaethusa Glöer & Manolas, 2024
Pikasia Taybi, Glöer & Mabrouki, 2021
† Pingyispira Y.-T. Li, 1986
Polatenia Georgiev, 2023
† Pomatiasia Guzhov, 2022
Pontohoratia Vinarski, Palatov & Glöer, 2014
†Pontohydrobia Badzoshvili, 1979
† Potamaclis F. Sandberger, 1873
† Potamolithoides W. B. Marshall & Bowles, 1932
Probythinella Thiele, 1928
†Prososthenia Neumayr, 1869
Pseudopaludinella Mabille, 1877 (taxon inquirendum)
†Pyrgulella Harzhauser, Neubauer & Kadolsky in Harzhauser et al., 2016
† Renistoma W. Yü, 1977
†Rhodopyrgula Willmann, 1981
Rifia Ghamizi, 2020
†Robicia Brusina, 1897
†Saccoia Brusina, 1893
Salaeniella Boeters, Quiñonero-Salgado & Ruiz-Cobo, 2019
†Salakosia Willmann, 1981
†Salalahia Kadolsky, Harzhauser & Neubauer in Harzhauser et al., 2016
†Scalimelania Wenz, 1939
Schapsugia Chertoprud, Palatov & Vinarski, 2021
Sellia Raincourt, 1884
Sheitanok Schütt & Şeşen, 1991
Sitnikovia Chertoprud, Palatov & Vinarski, 2020
Sivasi Şahin, Koca & Yildirim, 2012
†Socenia Jekelius, 1944
†Staja Brusina, 1897
†Stalioa Brusina, 1870
†Staliopsis Rzehak, 1893
Stygobium Grego & Glöer, 2019
Syrofontana Schütt, 1983
Tachira Chertoprud, Palatov & Vinarski, 2021
Tefennia Schütt & Yıldırım, 2003
Terranigra Radoman, 1978
Torosia Glöer & Georgiev, 2012
†Tournouerina Schlickum, 1971
Tridentalia Glöer & Hirschfelder, 2023
† Unicarinata Pan, 1982
Vilertia López-Soriano, Quiñonero-Salgado, A. A. Suárez, Rolán & Glöer, 2022
†Vrazia Brusina, 1897
† Wallaua R. Janssen, 2007
Wuconcha Kang, 1983 [citation needed]
†Xestopyrguloides Willmann, 1981
Znassela Taybi, Glöer & Mabrouki, 2023
Notes
References
MolluscaBase eds. (2024). MolluscaBase. Hydrobiidae W. Stimpson, 1865. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=120 on 2024-06-18
Ponder W. F.; Walker K. F. (January 2003). "From mound springs to mighty rivers: The conservation status of freshwater molluscs in Australia". Aquatic Ecosystem Health and Management. 6 (1): 19–28. doi:10.1080/14634980301482. S2CID 83920162.
Kabat, Alan R.; Herhsler, Robert (1993). "The Prosobranch Snail Family Hydrobiidae (Gastropoda: Rissooidea): Review of Classification and Supraspecific Taxa" (PDF). Smithsonian Contributions to Zoology. 547 (547): 1–94. doi:10.5479/si.00810282.547. Retrieved 13 February 2014.
Radoman, P. (1983). Hydrobioidea a superfamily of Prosobranchia (Gastropoda). I. Systematics. Serbian Academy of Sciences and Arts Monographs DXLVII, Department of Sciences 571: 1–256. book description.
Wilke T.; Davis G. M.; Falniowski A.; Giusti F.; Bodon M.; Szarowska M. (2001). "Molecular systematics of Hydrobiidae (Mollusca: Gastropoda: Rissooidea): testing monophyly and phylogenetic relationships". Proceedings of the Academy of Natural Sciences of Philadelphia. 151 (1): 1–21. doi:10.1635/0097-3157(2001)151[0001:MSOHMG]2.0.CO;2. ISSN 0097-3157. S2CID 86258304.
Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
Taylor D. W. (1966). The Veliger 9(2): 181, 199.
Brusina (1882). Bulletino della Società Malacologica Italiana 7(13–19): 230.
Thiele (1925). Handbuch der Zoologie 5(1): 80.
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