Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Vetigastropoda
Ordo: Lepetellida
Superfamilia: Fissurelloidea
Familia: Fissurellidae
Subfamilia: Emarginulinae
Genus: Emarginula
Subgenera: E. (Emarginula) – E. (Subzeidora)
Species: E. adriatica – E. alveolata – E. amitana – E. amyda – E. bajula – E. bonfittoi – E. capuliformis – E. choristes – E. christiaensi – E. compressa – E. compta – E. concinna – E. conica – E. costae – E. crassa – E. dentigera – E. devota – E. divae – E. elata – E. elongata – E. emarginella – E. faveola – E. fissura – E. fissurata – E. fragilis – E. fujitai – Emarginula gigantea E. guernei – E. hataii – E. hosoyai – E. huzardi – E. imaizumi – E. incisura – E. intervecta – E. japonica – E. kashimaensis – E. kawamurai – E. laevis – E. maculata – E. montrouzieri – E. multistriata – E. nordica – E. notata – E. octaviana – E. papillosa – E. paucipunctata – E. phrixodes – E. poppeorum – E. pulchreclathrata – E. pumila – E. punctata – E. punctulum – E. pustula – E. rosea – E. rugosa – E. sakyraii – E. serricostata – E. sicula – E. solidula – E. souverbiana – E. striatula – E. tenera – E. tenuis – E. teramachii – E. textilis – E. tosaensis – E. tuberculosa – E. vadum – E. variegata – E. velascoensis
Name
Emarginula Lamarck, 1801
References
Lamarck, J.-B. 1801. Systême des Animaux sans Vertèbres, ou Tableau général des classes, des orders at des genres de ces animaux; présentant leurs caractères essentiels et leur distribution, d'après la considération de leurs rapports naturels et de leur organisation, et suivant l'arrangement établi dans les galeries du Muséum d'Histoire Naturelle, parmi leurs dépouilles conservées; précédé du discours d'ouverture du Cours de Zoologie, donné dans le Muséum National d'Histoire Naturelle l'an 8 de la République. Paris: Detreville, viii + 432pp. BHL Google Books Open access Reference page.
Romani, L., Rolán, E., Simone, L.R.L. & Crocetta, F. 2017. Emarginula poppeorum and Acteon fasuloi: replacement names for the preoccupied taxa Emarginula gigantea Poppe, 2008 and Acteon elongatus Castellanos, Rolán & Bartolotta, 1987 (Mollusca: Gastropoda). Zootaxa 4300(1): 149–150. DOI: 10.11646/zootaxa.4300.1.11. Reference page.
Emarginula is a genus of small keyhole limpets, marine gastropod molluscs in the family Fissurellidae.[1]
Description
The oval shell is obliquely conical with a backward-pointing recurved apex. There is a deep incision in the anterior margin. A distinct anal fasciole extends upwards from this incision. The surface is latticed, sculptured with radial ribs. There is no septum inside.[2]
Radula
The radula has a dental formula of [many].1.4.1.4.1.[many],[3] with a wide, rectangular, acuspate rachidian tooth.[4] The various genera have a few dozen rows of teeth.[3]
Species
According to the World Register of Marine Species (WoRMS), species within the genus Emarginula include:[1]
Emarginula adamsiana Sowerby in Adams & Sowerby, 1863
Emarginula adriatica Costa O.G., 1829
Emarginula agulhasensis Thiele, 1925
Emarginula alba Watson, 1897
Emarginula amyda Shikama, 1962
Emarginula angusta McLean, 1970
Emarginula annielangleitae Poppe & Tagaro, 2020
Emarginula apolonia Pacaud, 2015
Emarginula balicasagensis Poppe & Tagaro, 2020
Emarginula bicancellata Montrouzier, 1860
Emarginula bonfittoi Smriglio & Mariottini, 2001
Emarginula boucheti Poppe & Tagaro, 2020
Emarginula camilla Melvill & Standen, 1903
Emarginula candida Adams, 1852
Emarginula choristes Dall, 1925
Emarginula christiaensi Piani, 1985
Emarginula circumalbum Poppe & Tagaro, 2020
Emarginula clypeus Adams, 1852
Emarginula colipanoae Poppe & Tagaro, 2020
Emarginula compta Habe, 1953
Emarginula concinna Adams, 1852
Emarginula connectens Thiele, 1915
Emarginula connelli Kilburn, 1978
Emarginula convexa Hedley, 1907
Emarginula costulata Deshayes, 1863
Emarginula crassa Sowerby, 1812
Emarginula crassicostata Sowerby in Adams & Sowerby, 1863
Emarginula curvamen Iredale, 1925
Emarginula cuvieri Audouin, 1827 (species inquirenda)
Emarginula decorata Deshayes, 1863
Emarginula delonguevilleae Poppe & Tagaro, 2020
Emarginula devota Thiele, 1915
Emarginula dictya McLean, 1970
Emarginula dilecta Adams, 1852
Emarginula divae (van Aartsen & Carrozza, 1995)
Emarginula dubia Schepman, 1908
† Emarginula espibosensis Lozouet, 1999
† Emarginula fenestrata Deshayes, 1861 †
Emarginula fenestrella Deshayes, 1863
Emarginula fissura (Linnaeus, 1758) - slit limpet
† Emarginula fossorensis Lozouet, 1999
Emarginula foveolata Schepman, 1908
Emarginula foveolata fugitai Habe, 1953
Emarginula fragilis Yokoyama, 1920
Emarginula fulginea Adams, 1851
Emarginula gabensis (Gabriel, 1961)
† Emarginula galeriformis Marwick, 1928
† Emarginula gigantea Poppe, 2008
Emarginula gustavi Poppe & Tagaro, 2020
Emarginula hataii Habe, 1953
Emarginula hawaiiensis Dall, 1895
† Emarginula haweraensis Powell, 1931
Emarginula hosoyai Habe, 1953
Emarginula huzardii (Payraudeau, 1826)
Emarginula icosisculpta Simone & Cunha, 2014
Emarginula imaizumi Dall, 1926
† Emarginula imbricata Millet, 1865
Emarginula imella Dall, 1926
Emarginula intervecta Locard, 1898: synonym of Emarginula fissura (Linnaeus, 1758)
Emarginula janae Cossignani, 2020
Emarginula japonica Sowerby in Adams & Sowerby, 1863
Emarginula jungcheni K.-Y. Lai, 2019
† Emarginula kaiparica Laws, 1939
Emarginula kashimaensis Shikama, 1962
† Emarginula komitica Laws, 1939
Emarginula koon Kilburn, 1978
Emarginula lata J.R.C. Quoy & J.P. Gaimard, 1834
Emarginula liuzzii Poppe & Tagaro, 2020
Emarginula longifissa Sowerby, 1866
Emarginula lorenzoi Giusti & Micali, 2019
Emarginula macclurgi Kilburn, 1978
Emarginula mactanensis Poppe & Tagaro, 2020
Emarginula maculata Adams in Adams & Sowerby, 1863
Emarginula margarita Poppe & Tagaro, 2020
Emarginula micans Adams, 1852
Emarginula multistriata Jeffreys, 1882
Emarginula natalensis Barnard, 1963
Emarginula nigromaculata (Thiele, 1930)
Emarginula nordica Pérez Farfante, 1947 - northern emarginula
† Emarginula oblonga Sandberger, 1859
Emarginula obovata Adams, 1852
Emarginula octaviana Coen, 1939
Emarginula oppressa Barnard, 1963
Emarginula paivana (Crosse, 1867)
Emarginula patula Cotton, 1930
† Emarginula paucicostata Laws, 1936
Emarginula paucipunctata Schepman, 1908
Emarginula philippinensis Poppe & Tagaro, 2020
Emarginula phrixodes Dall, 1927 - ruffled emarginula
Emarginula pittensis Marwick, 1928
Emarginula poppeorum Romani & Crocetta, 2017
† Emarginula praesicula Lozouet, 1999
Emarginula puncticulata Adams, 1852
Emarginula punctulum Piani, 1980
Emarginula pustula Thiele in Küster, 1913
Emarginula quadrata Poppe & Tagaro, 2020
Emarginula regia Habe, 1953
Emarginula retecosa Adams, 1851
Emarginula rosea Bell, 1824
Emarginula scabriuscula Adams, 1852
Emarginula sicula Gray, 1825 - dagger emarginula
Emarginula sinica Lu, 1986
Emarginula solidula Costa, 1829
Emarginula souverbiana Pilsbry, 1890
Emarginula spinosa Deshayes, 1863
Emarginula striatula Quoy and Gaimard, 1834 - dagger emarginula [5]
Emarginula subtilitexta Verco, 1908
Emarginula superba Hedley, 1906
Emarginula suspira Simone & Cunha, 2014
Emarginula tenera Locard, 1892
† Emarginula teneraeformis Lozouet, 1999
Emarginula tenuicostata Sowerby, 1863
Emarginula teramachii Habe, 1953
Emarginula textilis Gould, 1859 in 1859-61
Emarginula thomasi Crosse, 1864
Emarginula thorektes Kilburn, 1978
Emarginula tosaensis Habe, 1953
Emarginula tuberculosa Libassi, 1859 - tuberculate emarginula
Emarginula undulata Melvill & Standen, 1903
Emarginula velascoensis Shasky, 1961
Emarginula velascoi Rehder, 1980
Emarginula viridibrunneis Poppe & Tagaro, 2020
Emarginula viridicana Herbert & Kilburn, 1985
Emarginula viridis Poppe & Tagaro, 2020
Emarginula xishaensis Lu, 1986
Emarginula yangeorum Poppe & Tagaro, 2020
The following species are mentioned in other databases : The Indo-Pacific Molluscan Database also mentions the following species with names in current use :[6]
Emarginula curvata Schepman, 1908
Emarginula multisquamosa Schepman, 1908
Emarginula sublaevis Schepman, 1908
Subgenus Emarginula Lamarck, 1801
Emarginula bellula Adams, 1851
Emarginula cucullata Adams, 1851
Emarginula fissurata Adams, 1851
Emarginula harmilensis Sturany, 1903
Emarginula plana Schepman, 1908
Emarginula scabricostata Adams, 1851
Subgenus Emarginula (Subzeidora) Iredale, 1924
Species brought into synonymy
[1]
Emarginula altilis Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
Emarginula amitina Iredale, 1925: synonym of Emarginula dilecta A. Adams, 1852
Emarginula angustata Thiele, 1915: synonym of Tugali decussata A. Adams, 1852
Emarginula arabica A. Adams, 1852: synonym of Octomarginula arabica (A. Adams, 1852)
Emarginula arconatii Issel, 1869: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
Emarginula australis Quoy & Gaimard, 1834: synonym of Montfortia subemarginata (Blainville, 1819)
Emarginula bella Gabb, 1865: synonym of Scelidotoma bella (Gabb, 1865)
Emarginula biangulata Sowerby, 1901: synonym of Emarginella biangulata (G. B. Sowerby III, 1901)
† Emarginula blainvillii Defrance, 1825: synonym of † Rimula blainvillii (Defrance, 1825)
Emarginula brevirimata Deshayes, 1862: synonym of Montfortula brevirimata (Deshayes, 1862)
Emarginula clathrata A. Adams & Reeve, 1850: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
Emarginula clathrata Pease, 1863: synonym of Emarginula dilecta A. Adams, 1852
Emarginula conica Lamarck, 1801: synonym of Emarginula fissura (Linnaeus, 1758)
Emarginula conica Schumacher, 1817 non Lamarck, 1801: synonym of Emarginula rosea Bell, 1824
Emarginula dentigera Heilprin, 1889: synonym of Hemimarginula dentigera (Heilprin, 1889)
Emarginula elata Locard, 1898: synonym of Emarginula christiaensi Piani, 1985
Emarginula elongata O. G. Costa, 1829: synonym of Emarginula octaviana Coen, 1939
Emarginula eximia A. Adams, 1852: synonym of Emarginella eximia (A. Adams, 1852)
Emarginula flindersi Cotton, 1930: synonym of Emarginula convexa Hedley, 1907
Emarginula fuliginea Kuroda, 1941: synonym of Variemarginula variegata (A. Adams, 1852)
Emarginula guernei Dautzenberg & Fischer, 1896: synonym of Emarginula tuberculosa Libassi, 1859
Emarginula hedleyi Thiele, 1915: synonym of Emarginula candida A. Adams, 1852
Emarginula incisula(sic): synonym of Emarginella incisura (A. Adams, 1852)
Emarginula incisura A. Adams, 1852: synonym of Emarginella incisura (A. Adams, 1852)
Emarginula kimberti Cotton, 1930: synonym of Laeviemarginula kimberti (Cotton, 1930)
Emarginula modesta H. Adams, 1872: synonym of Hemimarginula modesta (H. Adams, 1872)
Emarginula muelleri Forbes & Hanley, 1849: synonym of Emarginula fissura (Linnaeus, 1758)
Emarginula ostheimerae Abbott, 1958: synonym of Octomarginula ostheimerae (Abbott, 1958)
Emarginula panhi Quoy & Gaimard, 1834: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
Emarginula papillosa Risso 1826: synonym of Emarginula huzardii Payraudeau, 1826
Emarginula parmophoidea Quoy & Gaimard, 1834: synonym of Tugali elegans Gray, 1843
Emarginula peasei Thiele, 1915: synonym of Emarginula dilecta A. Adams, 1852
Emarginula phrygium Herbert & Kilburn, 1985: synonym of Agariste phrygium (Herbert & Kilburn, 1986) (original combination)
Emarginula picta Dunker, 1860: synonym of Montfortula picta (Dunker, 1860)
Emarginula pileata Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
Emarginula pileum Heilprin, 1889: synonym of Hemimarginula pileum (Heilprin, 1889)
Emarginula planulata A. Adams, 1852: synonym of Emarginella planulata (A. Adams, 1852)
Emarginula polygonalis A. Adams, 1852: synonym of Hemitoma polygonalis (A. Adams, 1852)
Emarginula pulchreclathrata Tomlin, 1932: synonym of Emarginula agulhasensis Thiele, 1925
Emarginula pumila A. Adams, 1852: synonym of Hemimarginula pumila (A. Adams, 1852)
Emarginula punctata A. Adams, 1852: synonym of Variemarginula punctata (A. Adams, 1852)
Emarginula reticulata Sowerby, 1813: synonym of Emarginula fissura (Linnaeus, 1758)
Emarginula rugosa Quoy & Gaimard, 1834: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
Emarginula scutellata Deshayes, 1863: synonym of Octomarginula scutellata (Deshayes, 1863)
Emarginula sibogae Schepman, 1908: synonym of Emarginella sibogae (Schepman, 1908)
Emarginula simpla Christiaens, 1987: synonym of Hemimarginula simpla (Christiaens, 1987)
Emarginula stellata A. Adams, 1852: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
Emarginula subclathrata Pilsbry, 1890: synonym of Emarginula dilecta A. Adams, 1852
Emarginula subemarginata Blainville, 1819: synonym of Montfortia subemarginata (Blainville, 1819)
Emarginula subrugosa Thiele, 1916: synonym of Hemimarginula subrugosa (Thiele, 1916)
Emarginula sulcifera A. Adams, 1852: synonym of Montfortulana sulcifera (A. Adams, 1852)
Emarginula tricarinata Pilsbry, 1890: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
Emarginula tricostata G.B. Sowerby I, 1823: synonym of Hemitoma octoradiata (Gmelin, 1791)
Emarginula vadososinuata Yokoyama, 1922: synonym of Tugalina vadososinuata (Yokoyama, 1922)
Emarginula vadum Barnard, 1963: synonym of Emarginula undulata Melvill & Standen, 1903
Emarginula variegata A. Adams, 1852: synonym of Variemarginula variegata (A. Adams, 1852)
References
Bouchet, P.; Gofas, S.; Rosenberg, G. (2012). Emarginula Lamarck, 1801. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138012 on 2013-01-13
Tryon, George W. Jr. (1879–1898). Manual of Conchology, structural and systematic. Academy of Natural Sciences, Philadelphia. p. 248.
Torr, Claude M. (1914). "Radulæ of some South Australian Gasteropoda". Transactions of the Royal Society of South Australia. 38: 362–368.
Keen, A. Myra (1971-06-01). Sea shells of tropical west America: marine mollusks from Baja California to Peru. ISBN 978-0-8047-0736-7.
"Emarginula striatula". Gastropods.com. Retrieved 12 January 2019.
OBIS : Emarginula
"Emarginula". Integrated Taxonomic Information System.
Powell A W B, New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
Piani P. (1985). Revisione del genere Emarginula Lamarck, 1801 in Mediterraneo. Lavori, Società Italiana di Malacologia 21: 193-238
Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine Species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213
Rolán E., 2005. Malacological Fauna From The Cape Verde Archipelago. Part 1, Polyplacophora and Gastropoda.
Spencer, H.; Marshall. B. (2009). All Mollusca except Opisthobranchia. In: Gordon, D. (Ed.) (2009). New Zealand Inventory of Biodiversity. Volume One: Kingdom Animalia. 584 pp
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