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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
Superordo: Eupulmonata
Ordo: Stylommatophora
Subordo: Helicina
Infraordo: Limacoidei
Superfamilia: Limacoidea

Familia: Agriolimacidae
Subfamilia: Agriolimacinae - Mesolimacinae
Name

Agriolimacidae Wagner, 1935
References

Wagner, H. 1935. Diagnosen neuer Limaciden aus dem Naturhistorischen Museum in Wien. Zoologischer Anzeiger 95: 174.
Wiktor, A. 2000. Agriolimacidae (Gastropoda: Pulmonata) — a systematic monograph. Annales Zoologici (Warszawa) 49(3): 387.

Agriolimacidae is a family of small and medium-sized land slugs, or shell-less snails, terrestrial pulmonate gastropod mollusks.

Distribution

Distribution of Limacidae is Holarctic,[1] this include: Nearctic, western Palearctic and eastern Palearctic.[5]

Agriolimacidae is the largest slug family, some are introduced all over the world, synanthropes are often severe pests.[1]
This view of the right side of Deroceras praecox clearly shows the position of its pneumostome.
Anatomy

Most slugs in the family Agriolimacidae are rather small; only a few (in the genera Mesolimax and Krynickillus) are larger. Most are not more than 50 mm long.[1] The mantle is usually large, occupying approximately 1/3 of the entire body length, situated in the anterior part of the body.[1] The pneumostome is clearly postmedial.[1] The surface of the mantle in living slugs is covered in concentric, mobile wrinkles.[1] In addition sometimes there is a shallow, poorly defined groove which runs above the pneumostome on the right side, not passing to the left.[1]

The penis is short, usually bag-shaped, often with external appendages, inside with different stimulatory organs.[1] No tubular membrane encircles the penis and vas deferens.[1] The penis retractor muscle is situated beside the right tentacle.[1]

In this family, the number of haploid chromosomes lies between 26 and 30 (according to the values in this table).[6]
Taxonomy

The following two subfamilies have been recognized in the taxonomy of Bouchet & Rocroi (2005):

subfamily Agriolimacinae H. Wagner, 1935[2] - synonym: Deroceratinae Magne, 1952
subfamily Mesolimacinae Hausdorf, 1998

Genera

Genera within the family Agriolimacidae include:

subfamily Agriolimacinae

Deroceras Rafinesque, 1820 - type genus described as Agriolimax Mörch, 1865
Furcopenis Castillejo & Wiktor, 1983
Krynickillus Kaleniczenko, 1851

subfamily Mesolimacinae

Mesolimax Pollonera, 1888
Mesolimax brauni Pollonera, 1888

subfamily ?

Lytopelte Boettger, 1886
Megalopelte Lindholm, 1914

Cladogram

A cladogram showing the phylogenic relationships of this family to other families within the limacoid clade:[5]

limacoid clade 
 Staffordioidea 

Staffordiidae

 Dyakioidea 

Dyakiidae

 Gastrodontoidea 

Pristilomatidae

Chronidae

Euconulidae

Trochomorphidae

Gastrodontidae

Oxychilidae

 Parmacelloidea 

Trigonochlamydidae

Parmacellidae

Milacidae

 Zonitoidea 

Zonitidae

 Helicarionoidea 

Helicarionidae

Ariophantidae

Urocyclidae

 Limacoidea 

Vitrinidae

Boettgerillidae

Limacidae

Agriolimacidae


References

This article incorporates public domain text from the reference.[1]

"Family summary for Agriolimacidae". AnimalBase, last change 23-05-2010, accessed 4 August 2010
Wagner, J. (1935). "Magyarország, Horvátország és Dalmácia hazátlan csigái (40 szövegrajzzal). II. rész". Annales historico-naturales Musei nationalis hungarici. Pars Zoologica. 29: 169–212.
"Genera in family Agriolimacidae" [n = 6]. AnimalBase, accessed 10 September 2010.
Wiktor, A. (2000). "Agriolimacidae (Gastropoda: Pulmonata): a systematic monograph". Annales Zoologici 49: 347-590. abstract
Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.

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