Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Otomorpha
Subcohors: Ostariophysi
Sectio: Otophysa
Ordo: Cypriniformes
Subordo: Cyprinoidei
Familia: Cyprinidae
Subfamilia: Smiliogastrinae
Genus Systomus
Valid species currently placed in Systomus:
S. asoka – S. compressiformis – S. jacobusboehlkei – S. jayarami – S. martenstyni – S. orphoides – S. pleurotaenia – S. sarana – S. spilurus – S. timbiri
Remaining Southeast Asian Systomus sensu Rainboth (1996), filled up with more species from various other authors, belong to different lineages, e.g.:
S. aurotaeniatus
The "Puntius binotatus group" (including species of the "Lanao Puntius" and "Mindanao Puntius", as well as those from Lake Manguao on Palawan and Mindoro, plus additional species with similar characters) [Barbodes is available for the "Puntius binotatus group", with Cephalakompsus, Mandibularca, Ospatulus and Spratellicypris as synonyms]: P. amarus – S. banksi – S. binotatus – P. bantolanensis – P. cataractae – P. disa – S. dunckeri – S. everetti – P. flavifuscus – P. hemictenus – P. herrei – P. joaquinae – P. katolo – S. kuchingensis – P. lanaoensis – S. lateristriga – P. lindog – P. manalak – P. manguaoensis – P. rhombeus – P. sealei – P. semifasciolatus – P. sirang – P. tumba – P. umalii – S. xouthos
Dubious species and nom. nud. originally "described" as Systomus:
S. barbatus – S. hamiltonii – S. kadoon – S. siamensis – S. tripunctatus – S. unimaculatus
Species that were not included in the latest revision, however, were formerly placed in Systomus:
S. leiacanthus – S. takhoaensis
Name
Systomus McClelland, 1839
Type species: Systomus immaculatus McClelland, 1839
References
Primary references
McClelland, J. 1839. Indian Cyprinidae. Asiatic researches 19(2): 217–471.
Additional references
Kottelat, M. & Tan, H.H. 2011. Systomus xouthos, a new cyprinid fish from Borneo, and revalidation of Puntius pulcher (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters 22(3): 209–214. Reference page.
Parenti, L. R. & Lim, K. K. P. 2005. Fishes of the Rajang Basin, Sarawak, Malaysia. The Raffles Bulletin of Zoology, Suppl. no. 13: 175–208.
Pethiyagoda, R., Meegaskumbura, M. & Maduwage, K. 2012. A synopsis of the South Asian fishes referred to Puntius (Pisces: Cyprinidae). Ichthyological Exploration of Freshwaters 23(1): 69–95.
Rainboth, W.J. 1996. FAO species identification field guide for fishery purposes. Fishes of the Cambodian Mekong. Rome, FAO. 1–265. Reference page.
Links
Systomus and its species (including synonyms) in Catalog of Fishes, Eschmeyer, W.N., Fricke, R. & van der Laan, R. (eds.) 2024. Catalog of Fishes electronic version.
Systomus is a genus of fish in the family Cyprinidae native to tropical Asia.[1]
Species
There are currently 20 recognized species in this genus:[2]
Systomus asoka (Kottelat & Pethiyagoda, 1989) (Asoka barb)
Systomus binduchitra (Hora, 1937)
Systomus chryseus Plamoottil, 2014 [3] (Golden systomus)
Systomus chrysopoma (Valenciennes, 1842)
Systomus clavatus (McClelland, 1845) (Stedman barb)
Systomus compressiformis (Cockerell, 1913)
Systomus endecanalis (T. R. Roberts, 1989)
Systomus gracilus Plamoottil & Maji, 2020 (skinny systomus)
Systomus immaculatus McClelland, 1839
Systomus jacobusboehlkei Fowler, 1958
Systomus jayarami (Vishwanath & Tombi Singh, 1986)
Systomus laticeps Plamoottil, 2016 [4]
Systomus martenstyni (Kottelat & Pethiyagoda, 1991) (Martenstyn's barb)
Systomus orphoides (Valenciennes, 1842)
Systomus pleurotaenia (Bleeker, 1863) (Black lined barb)
Systomus rubripinnis (Valenciennes, 1842) (Javaen barb)
Systomus rufus Plamoottil, 2014 [3] (Red-finned kuruva)
Systomus sarana (F. Hamilton, 1822) (Olive barb)
Systomus spilurus (Günther, 1868)
Systomus subnasutus (Valenciennes, 1842)
Systomus timbiri (Deraniyagala, 1963) (Short-bodied olive barb)
References
Kottelat, M. (2013): The fishes of the inland waters of southeast Asia: A catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. Archived 2015-01-06 at the Wayback Machine Raffles Bulletin of Zoology, Suppl. No. 27: 1–663.
Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Systomus". FishBase. July 2024 version.
Plamoottil, M. (2014): Puntius nelsoni, Systomus chryseus and S. rufus (Cypriniformes: Cyprinidae), three new fish species from Kerala, India. International Journal of Fauna and Biological Studies, 1 (6): 135–145.
Plamoottil, M. (2016): Systomus laticeps, a new species of cyprinid fish from Kerala, India. International Journal of Fauna and Biological Studies, 3 (5): 92-96.
Plamoottil, M. & Maji, D, (2020) Systomus gracilus: a new fish (Cypriniformes: Cyprinidae) species from West Bengal. Journal of Experimental Zoology, India. 23 (2): 1033- 1038.
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