Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Otomorpha
Subcohors: Ostariophysi
Sectio: Otophysa
Ordo: Cypriniformes
Subordo: Cobitoidei
Familia: Nemacheilidae
Genus: Homatula
Species: H. acuticephala – H. anguillioides – H. anteridorsalis – H. change – H. coccinocola – H. cryptoclathrata – H. disparizona – H. erhaiensis – H. laxiclathra – H. longidorsalis – H. nanpanjiangensis – H. nigra – H. oligolepis – H. potanini – H. pycnolepis – H. robusta – H. variegata – H. wuliangensis
Name
Homatula Nichols, 1925: 2 [originally created as subgenus of Barbatula]
Type species: Nemachilus potanini Günther, 1896. Type by original designation (also monotypic).
Discussion
Synonym of Paracobitis Bleeker, 1863 (see: Zhu, 1989)
References
Endruweit, M., Min, R. & Yang, J-X. 2018. A new species of Homatula from the Red River drainage in Yunnan based on morphological and genetic data (Teleostei: Nemacheilidae). Zootaxa 4375(4): 555–566. DOI: 10.11646/zootaxa.4375.4.5 Reference page.
Kottelat, M. (1990): Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Verlag Dr. Friedrich Pfeil, München: 1–262.
Li, X., Che, X-J. & Zhou, W. 2019. Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species. Zootaxa 4711(2): 330–348. DOI: 10.11646/zootaxa.4711.2.6 Paywall Reference page.
Min, R., Yang, J.-X. & Chen, X.-Y. 2012. Homatula wuliangensis (Teleostei: Nemacheilidae), a new loach from Yunnan, China. Zootaxa 3586: 313–318. Preview PDF Reference page.
Min, R., Zhao, Y-H., Shi, J-S. & Yang, J-X. 2022. A new species of Homatula (Teleostei, Cobitoidea, Nemacheilidae) from the Pearl River drainage, Yunnan, China. Zookeys 1089ː 109–124. DOI: 10.3897/zookeys.1089.77203 Open access Reference page.
Nichols, J. T. (1925): Nemacheilus and related loaches in China. American Museum Novitates, 171: 1–7.
Zhu, S.-Q. (1989): The loaches of the subfamily Nemacheilinae in China (Cypriniformes: Cobitidae). Jiangsu Science & Technology Publishing House, Nanjing, China: i-x, 1-150. [In Chinese with English summary.]
Min, R., Chen, X.-Y., Yang, J.-X., Winterbottom, R. & Mayden, R.L. 2012. Phylogenetic relationships of the genus Homatula (Cypriniformes: Nemacheilidae), with special reference to the biogeographic history around the Yunnan-Guizhou Plateau. Zootaxa 3586: 78–94. Preview PDF Reference page.
Homatula is a genus of stone loaches endemic to China.
Species
There are currently 21 recognized species in this genus:[1][2]
Homatula acuticephala (W. Zhou & J. C. He, 1993)
Homatula anguillioides (S. Q. Zhu & S. H. Wang, 1985)
Homatula anteridorsalis Li, Che & Zhou, 2019
Homatula berezowskii (Günther, 1896)
Homatula change Endruweit, 2015 [3]
Homatula cryptoclathrata Li, Che & Zhou, 2019
Homatula disparizona R. Min, J. X. Yang & X. Y. Chen, 2013 [4]
Homatula erhaiensis (S. Q. Zhu & W. X. Cao, 1988)
Homatula guanheensis Zhou, Ma, Wang, Tang, Meng & Nie, 2021
Homatula laxiclathra J. H. Gu & E. Zhang, 2012 [5]
Homatula longidorsalis (J. X. Yang, Y. R. Chen & Kottelat, 1994)
Homatula nanpanjiangensis (R. Min, X. Y. Chen & J. X. Yang, 2010)
Homatula nigra Li, Che & Zhou, 2019
Homatula oligolepis (W. X. Cao & S. Q. Zhu, 1989)
Homatula potanini (Günther, 1896)
Homatula pycnolepis Y. T. Hu & E. Zhang, 2010
Homatula robusta Min, Zhao, Shi & Yang, 2022
Homatula variegata (Dabry de Thiersant, 1874)
Homatula wenshanensis C. Li, H. Yang, W. Li & T. Liu, 2017
Homatula wujiangensis R. H. Ding & Q. X. Deng, 1990
Homatula wuliangensis R. Min, J. X. Yang & X. Y. Chen, 2012 [6]
References
Froese, Rainer and Pauly, Daniel, eds. (2024). Species of Homatula in FishBase. February 2024 version.
Kottelat, M. (2012): Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). Archived 2013-02-11 at the Wayback Machine The Raffles Bulletin of Zoology, Suppl. No. 26: 1-199.
Endruweit, M. (2015): Homatula change, a new nemacheilid loach from the upper Black River basin in Yunnan, China (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 26 (1): 65-72.
Min, R., Yang, J.-X. & Chen, X.-Y. (2013): Homatula disparizona, a new species of loach from the Red River drainage in China (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 23 (4) [2012]: 351-355.
Gu, J.-H. & Zhang, E. (2012): Homatula laxiclathra (Teleostei: Balitoridae), a new species of nemacheiline loach from the Yellow River drainage in Shaanxi Province, northern China. Environmental Biology of Fishes, 94 (4): 591-599.
Min, R., Yang, J.-X. & Chen, X.-Y. (2012): Homatula wuliangensis (Teleostei: Nemacheilidae), a new loach from Yunnan, China. Zootaxa, 3586: 313–318.
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